The Crusaders first appear in the World War II title The Invaders and capture the crew of a crashed Germanbomber. Accepted by the British people, the team (Captain Wings, Dyna-Mite, Ghost Girl, the Spirit of '76, Thunder Fist, and Tommy Lightning) becomes the official protectors of the current king (George VI), thereby displacing the Americansuperhero team the Invaders. The Crusaders are revealed to be guided by a cab driver and apparent British spy called "Alfie", who can cancel their powers courtesy of a technological belt the character wears.
Dyna-Mite, who has no memory of his former life, becomes suspicious and spies on Alfie, learning that he is a Nazi agent and is planning to use the heroes to assassinate the king. Dyna-Mite warns the Invaders, and Alfie instigates a fight between the heroes and the Crusaders. The Nazi is killed in a car crash attempting to escape from the android Human Torch. The Crusaders, now powerless, disband, although Dyna-Mite—revealed to be Roger Aubrey, the close friend of Brian Falsworth, the brother of Invader Spitfire—remains trapped in a minute form.[1]
Aubrey is eventually restored to normal height and adopts Falsworth's former heroic identity as the Destroyer, now calling himself the Mighty Destroyer (as Falsworth eventually succeeded his father, James Montgomery Falsworth, as Union Jack II).[2]
At the same time that the Invaders were meeting the Crusaders in Marvel Comics, DC Comics' Freedom Fighters were also facing off against a team called the Crusaders,[4] with the DC version of the Crusaders based upon Marvel Comics' Invaders (several of the Crusaders that fought the Freedom Fighters were really "comic book fans" named "Lennie" (Len Wein), "Marvin" (Marv Wolfman), "Arch" (Archie Goodwin), and "Roy" (Roy Thomas), as shown in Freedom Fighters #9).