吸蟲綱
吸虫 (学名:Trematoda )是寄生虫 的一种,为扁形动物门 吸虫纲 动物的总称,也称为瓜仁虫 。一些吸虫也被称为二口虫 (拉丁語:Distoma)。其名由来是因为它的口吸盘和腹吸盘都被认为是口而致,拉丁语 中,Di代表“二”,stoma代表“口”。
概述
吸蟲是非常有名的一種寄生蟲 ,大部分為雌雄同體 [ 1] ,可自體受精;生活史中可能有一個或多個中間宿主 ,以魚類 、人體 等多種脊椎動物 為最終宿主;成蟲產卵發育成纖毛幼蟲 (Miracidium,亦作毛蚴 ),纖毛幼蟲入侵中間宿主後,增殖並發育成尾動幼蟲 (cercaride,亦作尾蚴 ),再被最終宿主食入前會發育成曩狀幼蟲 (metacerariae,亦作曩蚴 )[ 2] 。吸蟲的成蟲有口吸盤及腹吸盤,其卵有卵蓋 (operculum)[ 2] ,蟲體在人體中最久可活30年,由生食淡水魚而感染。貓、狗為一種名為保蟲的吸蟲宿主,成蟲寄生於膽道 ,大量感染時可在膽囊中發現蟲體。曩狀幼蟲存在於淡水魚 時,醃漬、煙燻或脫水皆不能將其殺死,輕微感染時無症狀,重度感染會造成膽道阻塞、膽囊炎 、結石 、肝膿痬 或急性胰臟炎 等,蟲卵約於感染後1個月後可於糞便中發現[ 3] 。
分類
吸蟲綱物種原來分為单殖目 (Monogena)、複殖目 (Digenea)、盾腹目 (Aspidogastrea,亦作盾盘目 或楯盤目 )三個目[ 1] 。單殖、複殖所指的是物種的雄性生殖系统的输精管 的数目(单殖1个、複殖2个);而盾腹目的命名則沿於其特征,在其腹口 吸盘周围有一個盾牌狀的衬垫。當時的單殖目可再細分為三個亞目[ 1] :259-260 ,但現時單殖目已被提升至綱級,成為單殖綱 。餘下的兩個目合計的物種,推算約 18,000[ 4] 到 24,000[ 5] 個物種。有文獻把這兩個目提升到亞綱的地位。
複殖目又名二生目 ,是吸蟲綱絕大多數物種所屬,之下再分為腹口亞目 (Gasterostomata,寄生於魚腸內)及前口亞目 (Prosostomata,有59科)[ 1] ,其物種泛稱複殖吸蟲(digenetic trematode)是以寄生人體為對象的吸蟲。複殖吸蟲的基本結構和發育過程大致上相同,種類相當繁多且形態各自相差頗大,生活史有性世代(寄生於軟體動物,如螺螄、蚌類的階段)與無性世代(寄生於脊椎動物體內階段),基本生活型態是以卵生且多種幼蟲(毛蚴 、胞蚴 、雷蚴 、尾蚴 、囊蚴 )之後在成長為成蟲[ 6] 。這些吸虫中以人类为宿主[ 1] 的种类,有肺吸虫(卫氏肺吸虫 )、肝吸虫(中華肝吸蟲 、牛羊肝吸蟲 )、肠道吸虫 、横川吸虫 、血吸虫 (日本血吸虫 )、布氏薑片蟲 (Fasciolopsis buski)等。
楯盤目只有一科四屬,直接發育,無需中間宿主。今提升至亞綱 級,是為楯吸蟲亞綱 或楯盤亞綱 。
本纲包括以下目:
目的地位未定的科:
目、科的地位未定的属:
Acanthocephala Diesing, 1858
Achillurbainia Dollfus, 1939
Adolescaria Sinitzin, 1911
Afrobilharzia Le Roux, 1958
Agamodistomum Stossich, 1892
Ahemiuris Chauhan, 1954
Allomicrophallus Yamaguti, 1971
Allopseudopiclidophora Yamaguti, 1965
Alloschistorchis Yamaguti, 1970
Allotelorchis Goodman, 1988
Amphibothrium Frey & Leuckart, 1847
Amphistomulum Brandes, 1892
Amphocleithrium Price & Romero, 1969
Anatirenicola Odening, 1962
Ankyrocotyle Vlassenko, 1928
Antepharyngeum Witenberg, 1925
Antodera Pigulevsky, 1952
Aototrema Bean-Knudsen, Uhazy & Wagner, 1987
Aspalacitrema Deblock & Rausch, 1965
Aspina Tripathi, 1956
Astomum Schlotthauber, 1860
Asymphylodoroides Yamaguti, 1971
Athesmioides Cribb & Spratt, 1992
Atrioglabrum Gracenea, Montoliu & Deblock, 1993
Atriospinosum Deblock, 1973
Australodiscus Sey, 1983
Axine Yamaguti, 1938
Baiohelmins Cribb & Pearson, 1988
Bashkinovitrema Skrjabin, 1944
Bathycola Fuhrmann, 1928
Baughbilharzia Chauhan, Srivastava & Chauhan, 1973
Beauchampia Yamaguti, 1963
Bengaliniella Deblock, Mukherjee & Ghosh, 1970
Bilateracotyloides Ramalingam, 1961
Bilorchis Mehra, 1937
Biplacunella Unnithan, 1971
Bispina Tripathi, 1956
Bistellavulva Unnithan, 1971
Bivulvostella Unnithan, 1971
Blairium Brooks & Holcman, 1993
Bothriophoron Stiles & Goldberger, 1910
Brachionus Schrank, 1776
Brachylecithoides Yamaguti, 1971
Brachysaccus Johnston, 1912
Bravotrema Groschaft, 1970
Bullulina Hammerschmidt, 1838
Buxifrons Fukui, 1929
Buxifrons Näsmark, 1937
Bychowskicotyle Unnithan, 1971
Calicotyle Nybelin, 1941
Calicotylides Nybelin, 1941
Canaania Travassos, 1944
Capronia Richard, 1966
Carangixenus Unnithan, 1962
Catoptroides Looss, 1902
Caudorchis Talbot, 1933
Cauliorchis Stiles & Goldberger, 1910
Cesartrema Travassos & Kohn, 1964
Ceylonotrema Gussev, 1963
Chamaeleophilus Yamaguti, 1971
Chaseostrigea Ukoli, 1967
Cheilostomum Diesing, 1850
Chelonella Beneden & Hesse, 1863
Chinhuta Lal, 1937
Chiropterotarbus Bay-Schmith, 1972
Chiropterotrema Velez & Thatcher, 1992
Chiroptodendrium Skarbilovitch, 1943
Choanocotyle Jue Sue & Platt, 1998
Choerocotyle Baer, 1959
Choerocotyloides Prudhoe, Yeh & Khalil, 1964
Choristogonoporus Stunkard, 1938
Cladorchis Wang & Zhou, 1989
Cochinocotyle Gupta & Gupta, 1971
Coelomotrema Angel, 1970
Coelomotrema aegyptiaca [ 7]
Coitocoecum Nicoll, 1915
Combesia Mas-Coma, Roset & Mantoliu, 1985
Conspicum Bhalerao, 1936
Coryphogonimus Cribb & Spratt, 1990
Cotylegaster Cobbold, 1877
Cryptocephalus Srivastava, 1944
Cryptostomum Nitzsch, 1832
Cychlorchis Lühe, 1908
Cyclostoma Otto, 1823
Cymbiforma Yamaguti, 1933
Dactylodiscus Kamegai, 1971
Dasyurotrema Cribb, 1992
Decacotylus Mayer, 1841
Dermophthiroides Cheung & Nigrelli, 1983
Diagenites Poinar & Boucot, 2006
Diastomum Packard, 1882
Dicotyle Sonsino, 1898
Dicranocercaria Lutz, 1919
Dicranocercaria Szidat, 1932
Digenites Poinar & Boucot, 2006
Dinotaster Unnithan, 1971
Diplobothrium Chandler, 1942
Diplostomum Brandes, 1888
Diplotrema Tripathi, 1959
Diporpa Dujardin, 1844
Distemum
Distoma Retzius, 1786
Distomulum Brandes, 1892
Distomum Diesing, 1850
Distomus Laennec, 1807
Divitelluses Cai & Li, 1993
Dolichodemas Looss, 1900
Dolichosomum Looss, 1899
Dozenkotrema Onkarnatha, 1964
Duttiella Srivastava, Prasad & Maurya, 1980
Echinella van Beneden & Hesse, 1864
Edcaballerotrema Teixeira de Freitas, 1960
Eroliostrigea Yamaguti, 1971
Eucercaria Diesing, 1855
Eucercarioides Tang & Tang, 1992
Eucradium Szidat, 1954
Eurobilharzia Le Roux, 1958
Eurymetra Odhner, 1910
Eustemma Diesing, 1850
Eustomos MacCallum, 1921
Evandrocotyle Jansen, 1941
Evanorchis Skrjabin, 1944
Exorchocoelium Thapar, 1958
Falciungus Akmerov, 1952
Festucaria Schrank, 1788
Flabellodiscus Johnston & Tiegs, 1922
Fontius Lotz & Palmieri, 1985
Gastrothylacias Stiles & Goldberger, 1910
Gastrothylacides Stiles & Goldberger, 1910
Geaterinia Lebedev, 1969
Gemellicotyle Prudhoe, 1975
Gibsonium Brooks & Holcman, 1993
Gigantatrium Yamaguti, 1958
Gigantobilharziella Khalifa, 1974
Gigantopora Looss, 1907
Glenocercaria Diesing, 1858
Globostomum Oshmarin, 1970
Globularia Rolando, 1841
Glossidioides Yamaguti, 1958
Glossocotyle Van Beneden & Hesse, 1863
Glyptamphistoma Yamaguti, 1958
Gotocotyla Meserve, 1938
Gymnocephala Diesing, 1858
Gymnocephala Leidy, 1877
Haematobium Dunglison, 1893
Haplostomum Burmeister, 1856
Harmostomum Witenberg, 1925
Hawkesius Stiles & Goldberger, 1910
Hemistephanus Looss, 1901
Hemixenotrema Baer, 1971
Hepatotrema Stunkard, 1923
Hephostomum Burmeister, 1856
Heptostomum Schomburgk, 1844
Heteracanthus Diesing, 1836
Heteraxine Linton, 1940
Heterobdella Beneden & Hesse, 1864
Heterocoelium Travassos, 1921
Heterocotyloides Yamaguti, 1963
Heterolope Looss, 1899
Heterostoma De Filippi, 1837
Hexastoma Kuhn, 1829
Hexastoma Rudolphi, 1809
Hexathyridium Treutler, 1793
Histerolecitha Linton, 1910
Histrionella Bory de St.Vincent, 1823
Histrionella Ehrenberg, 1828
Histrionellina Diesing, 1858
Holostoma Nitzsch, 1816
Hormocercaria Diesing, 1855
Hydricuculus McCrady, 1873
Hydrochoeristrema Sinkoc, Muller & Brum, 1998
Hypostoma Rudolphi, 1809
Indopleurogenes Yamaguti, 1971
Indopodocotyle Gupta & Puri, 1981
Indopodocotyloides Gupta & Sayal, 1979
Ishiella Unnithan, 1971
Ityogonismus
Jamuartrema Kumar Lal, 1976
Kaurma Chatterji, 1936
Khalilloossia Hilmy, 1949
Klepsitromis Hammerschmidt, 1835
Kunckelia Künstler, 1881
Labontes MacCallum, 1927
Labriferoides Ganapati, Hanumantha & Lakshmana Rao, 1962
Laterostrigea Yang, 1962
Leiperocotyle Eduardo, 1980
Leipertrema Sandosham, 1951
Lepidotes Johnston & Tiegs, 1922
Lepustomum Jain, 1953
Levinseniella Deblock & Pearson, 1971
Levinsensia Looss, 1928
Limatuloides Dubois, 1964
Limosina Manter, 1944
Lintonia Monticelli, 1904
Lobostoma Berthold, 1827
Longitrema Chen, 1954
Lophocercaria Diesing, 1858
Loriformis Goeze, 1782
Lutianicotyle Mamaev, 1990
Lutziella Rohde, 1966
Luxitrema Bray, 1987
Lyperosomoides Yamaguti, 1971
Macallumia Chatterji, 1938
Macrechinostomum Odhner, 1911
Macrocercus Müller, 1773
Macrocraeoides
Macrogenotrema Skrjabin & Baskakov, 1925
Macrurochaeta Costa, 1864
Maillardiella Brooks & Holcman, 1993
Malleolus Ehrenberg, 1838
Manidicola Graber & Gevrey, 1981
Maritremopsis Deblock, 1973
Maxtrema Travassos, Teixeira de Freitas & Kohn, 1969
Mazzantia Travassos & Kohn, 1964
Mediantestis Issaitschikow, 1928
Mediodera Pigulevsky, 1952
Mediopharyngeum Witenberg, 1925
Mediorchis Panin, 1984
Mediorima Nicoll, 1914
Meditypus Barton, 1994
Megacetabulum Oshmarin, 1964
Megacustis Bennett, 1935
Megalatriotrema Rao, 1969
Megalophalloides Ching & Ibanez Herrera, 1976
Mehraformis Bhardwaj, 1963
Melanoglena Eichwald, 1850
Mesodendrium Faust, 1919
Mesogonimus Monticelli, 1885
Metacercaria
Metadelphis Travassos, 1944
Metagorgoderina Fernandes, 1958
Metagyrodactylus Yamaguti, 1963
Metahelicometra Yamaguti, 1971
Microcotyle Tripathi, 1956
Microderma Mehra, 1931
Microphallus Deblock & Pearson, 1969
Mixophthalmus Karyakarte, 1967
Monarrhenos Deblock & Pearson, 1971
Monocaecum Deblock & Pearson, 1970
Monostoma Rudolphi, 1809
Monostoma Zeder, 1800
Monostomeus Rafinesque, 1815
Monostomulum Brandes, 1892
Morenodendrium Odening, 1969
Muhlingina Mehra, 1950
Multiglandularis Schulz & Skworzow, 1931
Multigonadus Bilqees, 1970
Multilecithum Oshmarin & Demshin, 1972
Musalia Shipley & Hornell, 1904
Muttua Shipley & Hornell, 1904
Nannoenteron Ozaki, 1924
Navigiolum Onji & Nishio, 1924
Neodactylodiscus Kamegai, 1972
Neodermophtherius Price, 1963
Neodicrocoelium Agrawal & Sharma, 1990
Neodiplostomum Dubois, 1937
Neoeuparadistomum Farooq & Azra Qamar, 1996
Neogorgoderina Pereira & Cuocolo, 1940
Neogyrodactylus Baugh, 1957
Neohamatopeduncularia Gupta & Varsha Sachdeva, 1987
Neolepoderma Mehra, 1937
Neomicrophallus Farooq & Yousuf, 1986
Neophroechinostoma Oshmarin & Belous, 1951
Neopisthorchis Chatterji & Kruidenier, 1961
Neoprosotocus Bourgat & Combes, 1977
Neorenifer Byrd & Denton, 1938
Nephrocephala Diesing, 1858
Neuronaia Goodsir, 1845
Nicollina Deshmukh, 1988
Nidarosia
Nidrosia Mortensen, 1933
Niviventertrema Dronen, Homesley & Cleveland, 1995
Notaster Unnithan, 1971
Octostoma Otto, 1823
Octothilus Hermann, 1782
Oligolecithus Vercammen-Grandjean, 1960
Olssoniella Travassos, 1944
Olveria Sinha, 1941
Olveria Thapar & Singha
Omnipharynx Nicholls & Brooks, 1969
Ompoktrema Varma & Singh, 1986
Onchiodiscus Kulkarni, 1969
Onchogaster Claparède, 1863
Ophiorchis Mehra, 1937
Opisthoarchiotrema Gupta & Sharma, 1974
Opisthobrachylecithum Yamaguti, 1971
Opisthogenes Nicoll, 1914
Opistholecithum Baer, 1970
Opistocotyle LeBlond, 1836
Oreochromogyrus Ferdousi & Chandra, 2002
Orientoacanthostomum Zaidi & Khan, 1977
Orthorchis Mödlinger, 1925
Orthorchis Travassos, 1944
Oryzomytrema Thatcher, 1982
Oswaldoia Travassos, 1919
Overstreetium Brooks & Holcman, 1993
Palamphistomum Srivastava & Tripathi, 1987
Palitrema Cogate, 1939
Pancreaticum Bhalerao, 1936
Pancreatrema Oshmarin, 1952
Papillatrium Richard, 1966
Papilloleucochloridium Bakke, 1980
Parabascoides Stunkard, 1938
Parabenedenia Johnston, 1929
Paracanthatrium Dubois, 1961
Paracephalogoniomus Skrjabin, 1949
Paraconcinnum Vassiliades & Richard, 1970
Paradistomoidella Brooks & Palmieri, 1981
Paradistomoides Travassos, 1944
Paraibatrema Ueta, Deberaldini, da Silva Cordeiro & de Toledo Artigas, 1981
Paraindopleurogenes Kalyankar & Palladwar, 1977
Paraleyogonimus Vaucher, 1968
Paralutztrema Faust, 1967
Parametadelphis Travassos, 1955
Paramonostomoides Yamaguti, 1971
Paramphistomacotyle Mehra, 1980
Paramphistomoides Yamaguti, 1958
Paraoistosomum Tkach, 2011
Paraplagiorchis Dollfus, 1924
Paraschistosomatium Price, 1929
Parepibdella Johnston, 1929
Paryphostomum Jain, 1952
Pentadion Saint-Rémy, 1891
Petalodistomum Pigulevsky, 1952
Philline Parona & Perugia, 1890
Phillines Westwood, 1840
Phrynobatrachotrema Goodman, 1986
Placotrema Looss, 1907
Plagiorchoides Olsen, 1937
Platyamphistoma Yamaguti, 1958
Platycotyle Beneden & Hesse, 1863
Platymetra Mehra, 1931
Platynosomoides Yamaguti, 1971
Plenosominoides Ke & Liang, 1984
Pleurolobatus Kaw, 1944
Pleuropsoloides Goodman, 1990
Polyporus Grube, 1840
Postodera Pigulevsky, 1952
Postpharyngeum Witenberg, 1925
Praeorchitrema Oshmarin, 1952
Primatotrema Premvati, 1958
Proacetabulorchoides Oshmarin, 1970
Probolitrema Pigulevsky, 1952
Prochiorchis Dollfus, 1963
Procyonophilus Thatcher, 1982
Proechimytrema Thatcher, 1982
Proeputiodiplostomum Gupta, 1962
Progonimodiscus Vercammen-Grandjean, 1960
Promicrophallus Fischthal & Kuntz, 1981
Pronopharynx Cohn, 1904
Proschistosoma Grétillat, 1962
Proshystera Korkhaus, 1930
Prosostephanus Lutz, 1935
Prosthometra Looss, 1896
Prostorchigenes Tubangui, 1928
Protofasciola Odhner, 1926
Protofasciolopsis Odhner
Prouterina Foreyt, Schell & Beyer, 1996
Prudhoella Beverley-Burton, 1960
Prymnoprion Looss, 1899
Pseudalassostoma Yamaguti, 1958
Pseudallassostomoides Yamaguti, 1971
Pseudallobenedenia Yamaguti, 1966
Pseudathesmia Travassos, 1942
Pseudoallocreadium Yamaguti, 1971
Pseudobianium Sey, 1996
Pseudocalceostoma Yamaguti, 1963
Pseudodiplodiscoides Murty, 1970
Pseudodiscus Sonsino, 1895
Pseudoechinostomum
Pseudoganeo Khomenovskij, 1971
Pseudomagnivitellinum Dronen & Underwood, 1980
Pseudomerizocotyle Kay, 1942
Pseudomorcheides Wang, 1982
Pseudoorientocreadium Agrawal & Sharma, 1990
Pseudoparamphistoma Yamaguti, 1958
Pseudoplagiorchis Yamaguti, 1971
Pseudoquinqueserialis Sutton, 1981
Pseudorchis Yamaguti, 1971
Pseudorenifer Price, 1935
Pseudospelotrematoides Yamaguti, 1939
Pterocotyle Beneden & Hesse, 1863
Pteronella Beneden & Hesse, 1863
Pterygotomaschalos Stunkard, 1924
Ptyasiorchis Mehra, 1937
Quasimaritrema Deblock, 1973
Rallitrema Travassos & Kohn, 1964
Redia Filippi, 1837
Reduxotrema Pigulevsky, 1952
Renicola Sudarikov, 1951
Rhacophotrema Uchida, Itagaki & Inoue, 1980
Rhodobilharzia Le Roux, 1958
Rhopalocerca Diesing, 1850
Rudolphiella Travassos, 1924
Rutschurutrema Baer, 1959
Sarumitrema Beverley-Burton, 1963
Sawqirahcotyle Lebedev, 1976
Schisturus Rudolphi, 1809
Schizocerca Diesing, 1858
Scintillovulva Unnithan, 1971
Serpentinotrema Travassos & Kohn, 1964
Sigmapera Nicoll, 1918
华毕属 Sinobilharzia Dutt & Srivastava, 1955
Sinobilharzia Le Roux, 1958
Sinuosoides Denton & Krissinger, 1975
Skrjabinodendrium Skarbilovitch, 1943
Skrjabinosomum Skrjabin, 1944
Solenocerca Diesing, 1855
Solenocotyle Diesing, 1850
Sorexegliphe Sadovskaya, 1954
Spinoderetrema Bray, 1987
Spinometra Mehra, 1931
Sporocerca Diesing, 1855
Sporocystis Filippi, 1854
Sporocystophora Diesing, 1858
Sporonema Diesing, 1855
Sporotherium Diesing, 1858
Srivastavaia Singh, 1970
Staffordiella Mehra, 1966
Stephanolecithus Nakagawa, 1919
Stephanopharynx Fischoeder, 1901
Stichocotyle Faust & Tang, 1936
Strzeleckia Cribb & Spratt, 1991
Sturniratrema Velez & Thatcher, 1992
Suchocyathocotyle Dubois, 1984
Sureshiella Srivastava, Maurya & Prasad, 1980
Symmetricatesticula Krasnolobova, 1976
Synethes Stiles & Goldberger, 1910
Tagumaea Fukui, 1930
Taiwantrema Fischthal & Kuntz, 1981
Telogonella Mehra & Negi, 1928
Tenuifasciola Yamaguti, 1971
Testisaculus Bhalerao, 1927
Tetrapapillatrema Ralph, 1938
Tetrastoma Chiaje, 1833
Thailandobilharzia Baugh, 1977
Thylogaster Stiles & Goldberger, 1910
Topsiturvitrema Velez & Thatcher, 1992
Travassasella Faust & Tang, 1938
Travassodendrium Skarbilovitch, 1943
Travassosina Vaz, 1932
Travassostrema Rebecq, 1962
Triloborchidiplostomum Dubois & Angel, 1972
Trionchus MacCallum, 1916
Tumaclinostomum Van der Kuyp, 1953
Turbinilla Bory de St.Vincent, 1823
Turquinia Coy Otero, 1989
Ugandalubens Goodman, 1988
Ultragenotrema Skrjabin & Baskakow, 1925
Unistriata Filimonova, 1985
Vaginaespina Tripathi, 1956
Velasquezotrema Eduardo & Javellana, 1987
Verodunia Lahille & Joan, 1918
Vesperugidendrium Pande, 1937
Virgulina Bory de St.-Vincent, 1823
Vitellariotrema Gupta & Krishna, 1977
Voelkeria Travassos & Kohn, 1966
Watsonius Stiles & Goldberger, 1910
Wellmanius Stiles & Goldberger, 1910
Wrightrenicola Odening, 1962
Xenopharynx Nicoll, 1912
Xenopodistomum Macnae, Rock & Makowski, 1973
Xiphidiocercaria Diesing, 1855
Zeferinella Artigas & Paulino, 1988
Zeylanurotrema Crusz & Sanmugasunderam, 1974
Zoosperma Bory de St Vincent, 1823
Zygodendrium Salem, 1976
構造
體壁組織可分為體被與肌肉;可自我更新;具有吸收營養、感覺等功能[ 6] 。
参考文献
^ 1.0 1.1 1.2 1.3 1.4 李永基 . 第貳部無脊椎動物 第VII篇扁形動物門. 中山自然科學大辭典 . 第九冊:動物學. : 249, 259-272 [2016-09-23 ] . (原始内容存档 于2020-07-05) (中文(繁體)) .
^ 2.0 2.1 第二篇 蠕虫学:第六章 吸 虫:吸虫概论 . 人体寄生虫学 . 郑州大学 远程教育学院. [2016-09-25 ] . (原始内容 存档于2016-09-27) (中文(简体)) .
^ 溫惠敏 . 《石門水庫活魚3吃 陷危機》 . 壹週刊 (台灣) . 2011-02-03, 506 : 69 [2016-09-25 ] . (原始内容存档 于2019-05-15) (中文(繁體)) .
^ Littlewood D T J; Bray R. A. The Digenea. Interrelationships of the Platyhelminthes. Systematics Association Special Volume 60 1. CRC. 2000: 168–185. ISBN 978-0-7484-0903-7 .
^ Poulin, Robert; Serge Morand. Parasite Biodiversity. Smithsonian. 2005: 216. ISBN 978-1-58834-170-9 .
^ 6.0 6.1 吸虫概论 . 山东大学 . [2016-09-25 ] . (原始内容 存档于2011-05-14) (中文(简体)) .
^ Arlene Jones & O.M.E. El-Azazy (1986) Coelomotrema aegyptiaca sp. nov., an unusual prosthogonimid trematode from Rattus norvegicus (Berkenhout) in Egypt, Journal of Natural History, 20:3, 707-712, DOI: 10.1080/00222938600770481
参见