Hydrophiinae
Morske zmije , ili zmije koralnog grebena , su potporodica otrovnih elapidnih zmija , Hydrophiinae , koje nastanjuju morske sredine tokom većeg dela ili celokupnog života. Većina njih je dobro prilagođena potpuno vodenom životu i ne mogu se kretati kopnom, osim roda Laticauda , koji ima ograničeno kretanje kopnom. One se nalaze se u toplim obalskim vodama od Indijskog do Tihog okeana i usko su povezane sa otrovnim kopnenim zmijama u Australiji.[ 2]
Sve morske zmije imaju repove nalik veslu, a mnoge imaju bočno komprimovana tela koja im daju izgled jegulja . Za razliku od riba, nemaju škrge i moraju redovno da izrone da bi disale. Zajedno s kitovima , spadaju u najpotpunije vodene životinje od svih kičmenjaka koji dišu vazduh.[ 3] Neke od vrsta ove grupe imaju najmoćnije otrove od svih zmija. Deo njih ima blage dispozicije i ugrizaju samo kad su provocirane, dok su druge puno agresivnije. Trenutno je 17 rodova morskih zmija opisano.[ 4] [ 5] [ 5]
Sistematika
Rodovi
Acanthophis Daudin, 1803
Aipysurus Lacépède, 1804
Antaioserpens Wells and Wellington, 1985
Aspidomorphus Fitzinger, 1843
Austrelaps Worrell, 1963
Brachyurophis Günther, 1863
Cacophis Günther, 1863
Cryptophis Worrell, 1961
Demansia Günther, 1858
Denisonia Krefft, 1869
Drysdalia Worrell, 1961
Echiopsis Fitzinger, 1843
Elapognathus Boulenger, 1896
Emydocephalus Krefft, 1869
Ephalophis M. A. Smith, 1931
Furina A. M. C. Duméril, 1853
Hemiaspis Fitzinger, 1861
Hoplocephalus Wagler, 1830
Hydrelaps Boulenger, 1896
Hydrophis Latreille in Sonnini and Latreille, 1801
Kolpophis M. A. Smith, 1926
Loveridgelaps McDowell, 1970
Micropechis Boulenger, 1896
Neelaps Günther, 1863
Notechis Boulenger, 1896
Ogmodon W. C. H. Peters, 1864
Oxyuranus Kinghorn, 1923
Parahydrophis Burger and Natsuno, 1974
Parapistocalamus Roux, 1934
Parasuta Worrell, 1961
Paroplocephalus Keogh, Scott and Scanlon, 2000
Pseudechis Wagler, 1830
Pseudonaja Günther, 1858
Rhinoplocephalus F. Müller, 1885
Salomonelaps McDowell, 1970
Simoselaps Jan, 1859
Suta Worrell, 1961
Thalassophis P. Schmidt, 1852
Toxicocalamus Boulenger, 1896
Tropidechis Günther, 1863
Vermicella Gray in Günther, 1858
Vrste
Acanthophis antarcticus (Shaw and Nodder, 1802)
Acanthophis cryptamydros Maddodk, Ellis, Doughty, A. Smith and Wüster, 2015
Acanthophis hawkei Wells and Wellington, 1985
Acanthophis laevis Macleay, 1878
Acanthophis praelongus Ramsay, 1877
Acanthophis pyrrhus Boulenger, 1898
Acanthophis rugosus Loveridge, 1948
Acanthophis wellsi Hoser, 1998
Aipysurus apraefrontalis M. A. Smith, 1926
Aipysurus duboisii Bavay, 1869
Aipysurus eydouxii (Gray, 1849)
Aipysurus foliosquama M. A. Smith, 1926
Aipysurus fuscus (Tschudi, 1837)
Aipysurus laevis Lacépède, 1804
Aipysurus mosaicus Sanders, Rasmussen, Elmburg, Mumpuni, Guinea, Blias, Lee and Fry, 2012
Aipysurus pooleorum L. A. Smith, 1974
Aipysurus tenuis Lönnberg and Andersson, 1913
Antaioserpens albiceps (Boulenger, 1898)
Antaioserpens warro (De Vis, 1884)
Aspidomorphus lineaticollis (Werner, 1903)
Aspidomorphus muelleri (Schlegel, 1837)
Aspidomorphus schlegelii (Günther, 1872)
Austrelaps labialis (Jan, 1859)
Austrelaps ramsayi (Krefft, 1864)
Austrelaps superbus (Günther, 1858)
Brachyurophis approximans (Glauert, 1954)
Brachyurophis australis (Krefft, 1864)
Brachyurophis campbelli (Kinghorn, 1929)
Brachyurophis fasciolatus (Günther, 1872)
Brachyurophis incinctus (Storr, 1968)
Brachyurophis morrisi (Horner, 1998)
Brachyurophis roperi (Kinghorn, 1931)
Brachyurophis semifasciatus Günther, 1863
Cacophis churchilli Wells and Wellington, 1985
Cacophis harriettae Krefft, 1869
Cacophis krefftii Günther, 1863
Cacophis squamulosus (A. M. C. Duméril, Bibron and A. H. A. Duméril, 1854)
Cryptophis boschmai (Brongersma and Knaap-van Meeuwen, 1964)
Cryptophis incredibilis (Wells and Wellington, 1985)
Cryptophis nigrescens (Günther, 1862)
Cryptophis nigrostriatus (Krefft, 1864)
Cryptophis pallidiceps (Günther, 1858)
Demansia angusticeps (Macleay, 1888)
Demansia calodera Storr, 1978
Demansia flagellatio Wells and Wellington, 1985
Demansia olivacea (Gray, 1842)
Demansia papuensis (Macleay, 1877)
Demansia psammophis (Schlegel, 1837)
Demansia quaesitor Shea in Shea and Scanlon, 2007
Demansia reticulata (Gray, 1842)
Demansia rimicola Scanlon in Shea and Scanlon, 2007
Demansia rufescens Storr, 1978
Demansia shinei Shea in Shea and Scanlon, 2007
Demansia simplex Storr, 1978
Demansia torquata (Günther, 1862)
Demansia vestigiata (De Vis, 1884)
Denisonia devisi Waite and Longman, 1920
Denisonia maculata (Steindachner, 1867)
Drysdalia coronoides (Günther, 1858)
Drysdalia mastersii (Krefft, 1866)
Drysdalia rhodogaster (Jan in Jan and Sordelli, 1873)
Echiopsis curta (Schlegel, 1837)
Elapognathus coronatus (Schlegel, 1837)
Elapognathus minor (Günther, 1863)
Emydocephalus annulatus Krefft, 1869
Emydocephalus ijimae Stejneger, 1898
Emydocephalus szczerbaki Dotsenko, 2011
Ephalophis greyae M. A. Smith, 1931
Furina barnardi (Kinghorn, 1939)
Furina diadema (Schlegel, 1837)
Furina dunmalli (Worrell, 1955)
Furina ornata (Gray, 1842)
Furina tristis (Günther, 1858)
Hemiaspis damelii (Günther, 1876)
Hemiaspis signata (Jan, 1859)
Hoplocephalus bitorquatus (Jan, 1859)
Hoplocephalus bungaroides (Schlegel, 1837)
Hoplocephalus stephensii Krefft, 1869
Hydrelaps darwiniensis Boulenger, 1896
Hydrophis atriceps Günther, 1864
Hydrophis belcheri (Gray, 1849)
Hydrophis bituberculatus W. C. H. Peters, 1873
Hydrophis brookii Günther, 1872
Hydrophis caerulescens (Shaw, 1802)
Hydrophis cantoris Günther, 1864
Hydrophis coggeri (Kharin, 1984)
Hydrophis curtus (Shaw, 1802)
Hydrophis cyanocinctus Daudin, 1803
Hydrophis czeblukovi (Kharin, 1984)
Hydrophis donaldi Ukuwela, Sanders and Fry, 2012
Hydrophis elegans (Gray, 1842)
Hydrophis fasciatus (Schneider, 1799)
Hydrophis gracilis (Shaw, 1802)
Hydrophis hardwickii (Gray, 1835)
Hydrophis hendersoni (Boulenger, 1903)
Hydrophis inornatus (Gray, 1849)
Hydrophis jerdonii (Gray, 1849)
Hydrophis kingii Boulenger, 1896
Hydrophis klossi Boulenger, 1912
Hydrophis laboutei Rasmussen and Ineich, 2000
Hydrophis lamberti M. A. Smith, 1917
Hydrophis lapemoides (Gray, 1849)
Hydrophis macdowelli Kharin, 1983
Hydrophis major (Shaw, 1802)
Hydrophis mamillaris (Daudin, 1803)
Hydrophis melanocephalus Gray, 1849
Hydrophis melanosoma Günther, 1864
Hydrophis nigrocinctus Daudin, 1803
Hydrophis obscurus Daudin, 1803
Hydrophis ocellatus Gray, 1849
Hydrophis ornatus (Gray, 1842)
Hydrophis pachycercos Fischer, 1855
Hydrophis pacificus Boulenger, 1896
Hydrophis parviceps M. A. Smith, 1935
Hydrophis peronii (A. M. C. Duméril, 1853)
Hydrophis platurus (Linnaeus, 1766)
Hydrophis schistosus Daudin, 1803
Hydrophis semperi Garman, 1881
Hydrophis sibauensis Rasmussen, Auliya and Böhme, 2001
Hydrophis spiralis (Shaw, 1802)
Hydrophis stokesii (Gray, 1846)
Hydrophis stricticollis Günther, 1864
Hydrophis torquatus Günther, 1864
Hydrophis viperinus (P. Schmidt, 1852)
Hydrophis vorisi Kharin, 1984
Hydrophis zweifeli (Kharin, 1985)
Kolpophis annandalei (Laidlaw, 1901)
Loveridgelaps elapoides (Boulenger, 1890)
Micropechis ikaheca (Lesson in Duperrey, 1829)
Neelaps calonotus (A. M. C. Duméril, Bibron and A. H. A. Duméril, 1854)
Notechis scutatus (W. C. H. Peters, 1861)
Ogmodon vitianus W. C. H. Peters, 1864
Oxyuranus microlepidotus (McCoy, 1879)
Oxyuranus scutellatus (W. C. H. Peters, 1867)
Oxyuranus temporalis Doughty, Maryan, Donnellan and Hutchinson, 2007
Parahydrophis mertoni (Roux, 1910)
Parapistocalamus hedigeri Roux, 1934
Parasuta dwyeri (Worrell, 1956)
Parasuta flagellum (McCoy, 1878)
Parasuta gouldii (Gray, 1841)
Parasuta monachus (Storr, 1964)
Parasuta nigriceps (Günther, 1863)
Parasuta spectabilis (Krefft, 1869)
Paroplocephalus atriceps (Storr, 1980)
Pseudechis australis (Gray, 1842)
Pseudechis butleri L. A. Smith, 1982
Pseudechis colletti Boulenger, 1902
Pseudechis guttatus De Vis, 1905
Pseudechis pailsei (Hoser, 1998)
Pseudechis papuanus W. C. H. Peters and Doria, 1878
Pseudechis porphyriacus (Shaw, 1794)
Pseudechis rossignolii (Hoser, 2000)
Pseudechis weigeli (Wells and Wellington, 1987)
Pseudonaja affinis Günther, 1872
Pseudonaja aspidorhyncha (McCoy, 1879)
Pseudonaja guttata (Parker, 1926)
Pseudonaja inframacula (Waite, 1925)
Pseudonaja ingrami (Boulenger, 1908)
Pseudonaja mengdeni Wells and Wellington, 1985
Pseudonaja modesta (Günther, 1872)
Pseudonaja nuchalis Günther, 1858
Pseudonaja textilis (A. M. C. Duméril, Bibron and A. H. A. Duméril, 1854)
Rhinoplocephalus bicolor F. Müller, 1885
Salomonelaps par (Boulenger, 1884)
Simoselaps anomalus (Sternfeld, 1919)
Simoselaps bertholdi (Jan, 1859)
Simoselaps bimaculatus (A. M. C. Duméril, Bibron and A. H. A. Duméril, 1854)
Simoselaps littoralis (Storr, 1968)
Simoselaps minimus (Worrell, 1960)
Suta fasciata (Rosén, 1905)
Suta ordensis (Storr, 1984)
Suta punctata (Boulenger, 1896)
Suta suta (W. C. H. Peters, 1863)
Thalassophis anomalus P. Schmidt, 1852
Toxicocalamus buergersi (Sternfeld, 1913)
Toxicocalamus ernstmayri O'Shea, Parker and Kaiser, 2015
Toxicocalamus grandis (Boulenger, 1914)
Toxicocalamus holopelturus McDowell, 1969
Toxicocalamus longissimus Boulenger, 1896
Toxicocalamus loriae (Boulenger, 1898)
Toxicocalamus mintoni Kraus, 2009
Toxicocalamus misimae McDowell, 1969
Toxicocalamus pachysomus Kraus, 2009
Toxicocalamus preussi (Sternfeld, 1913)
Toxicocalamus spilolepidotus McDowell, 1969
Toxicocalamus stanleyanus Boulenger, 1903
Tropidechis carinatus (Krefft, 1863)
Vermicella annulata (Gray in Gray, 1841)
Vermicella intermedia Keogh and S. A. Smith, 1996
Vermicella multifasciata (Longman, 1915)
Vermicella snelli Storr, 1968
Vermicella vermiformis Keogh and S. A. Smith, 1996
Podvrste
Acanthophis antarcticus antarcticus (Shaw and Nodder, 1802)
Acanthophis antarcticus schistos Wells and Wellington, 1985
Acanthophis wellsi donnellani Hoser, 2002
Acanthophis wellsi wellsi Hoser, 1998
Aspidomorphus muelleri interruptus Brongersma, 1934
Aspidomorphus muelleri lineatus Brongersma, 1934
Aspidomorphus muelleri muelleri (Schlegel, 1837)
Brachyurophis fasciolatus fasciatus (Stirling and Zietz, 1893)
Brachyurophis fasciolatus fasciolatus (Günther, 1872)
Brachyurophis semifasciatus semifasciatus Günther, 1863
Brachyurophis semifasciatus woodjonesii (Thomson, 1934)
Demansia papuensis melaena Storr, 1978
Demansia papuensis papuensis (Macleay, 1877)
Demansia psammophis cupreiceps Storr, 1978
Demansia psammophis psammophis (Schlegel, 1837)
Hydrophis caerulescens caerulescens (Shaw, 1802)
Hydrophis caerulescens hybridus Schlegel, 1844
Hydrophis jerdonii jerdonii (Gray, 1849)
Hydrophis jerdonii siamensis (M. A. Smith, 1926)
Hydrophis ornatus godeffroyi W. C. H. Peters, 1873
Hydrophis ornatus maresinensis Mittleman, 1947
Hydrophis ornatus ornatus (Gray, 1842)
Hydrophis torquatus aagaardi M. A. Smith, 1920
Hydrophis torquatus diadema Günther, 1864
Hydrophis torquatus torquatus Günther, 1864
Micropechis ikaheca fasciatus (Fischer, 1884)
Micropechis ikaheca ikaheca (Lesson in Duperrey, 1829)
Notechis scutatus occidentalis Glauert, 1948
Notechis scutatus scutatus (W. C. H. Peters, 1861)
Oxyuranus scutellatus canni Slater, 1956
Oxyuranus scutellatus scutellatus (W. C. H. Peters, 1867)
Parasuta spectabilis bushi (Storr, 1988)
Parasuta spectabilis nullarbor (Storr, 1981)
Parasuta spectabilis spectabilis (Krefft, 1869)
Pseudonaja affinis affinis Günther, 1872
Pseudonaja affinis exilis Storr, 1989
Pseudonaja affinis tanneri Worrell, 1961
Pseudonaja textilis pughi Hoser, 2003
Pseudonaja textilis textilis (A. M. C. Duméril, Bibron and A. H. A. Duméril, 1854)
Toxicocalamus preussi angusticinctus (Bogert and Matalas, 1945)
Toxicocalamus preussi preussi (Sternfeld, 1913)
Reference
Literatura
Graham JB, Lowell WR, Rubinoff I, Motta J. 1987. Surface and subsurface swimming of the sea snake Pelamis platurus . J. exp. Biol. 127, 27-44. PDF at the [Journal of Experimental Biology]. Accessed 7 August 2007.
Rasmussen AR. 1997. Systematics of sea snakes; a critical review. Symp. Zool. Soc. London 70, 15-30.
Smith MA. 1926. Monograph of the sea snakes (Hydrophiidae). British Museum of Natural History, London.
Voris HK. 1977. A phylogeny of the sea snakes (Hydrophiidae). Fieldiana Zool. 70, 79-169.
Whitaker R. 1978. Common Indian Snakes: A Field Guide. Macmillan India Limited.
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