Simbirskiasaurus is an extinct genus of ichthyosaur from the Early Cretaceous (early Barremian) of Ulyanovsk Oblast, Russia.[1][2] Its type specimen is YKM 65119, a fragmentary skull and vertebral column.[3]
Fischer et al. give the diagnosis of Simbirskiasaurus as follows: "Platypterygiine ophthalmosaurid characterized by the following autapomorphies: external naris divided by a nasomaxillary pillar; posterior opening of the narial complex with anteroposteriorly constricted dorsal extension; deeply interdigitating prefrontal–lacrimal suture [reminiscent of the basal neoichthyosaurian Temnodontosaurus platyodon (Conybeare, 1822); see Godefroit, 1993]. Simbirskiasaurus birjukovi is also characterized by the following unique combination of features: subnarial process of the premaxilla reaches the posterior margin of the external naris (shared with Cryopterygius kristiansenae Druckenmiller et al., 2012); elongated anterior process of the maxilla, reaching anteriorly the level of the nasal [unlike in Aegirosaurus leptospondylus Bardet & Fernández, 2000 and Sveltonectes insolitus (Fischer et al., 2011b)]; pres- ence of a supranarial process of the premaxilla [shared with Platypterygius australis (McCoy, 1867), see Kear, 2005, and possibly Pervushovisaurus bannovkensis (Arkhangelsky, 1998b)]."[2]
The following cladogram shows a possible phylogenetic position of Simbirskiasaurus in Ophthalmosauridae according to the analysis performed by Zverkov and Jacobs (2020).[4]
Acamptonectes densus
Mollesaurus periallus
Ophthalmosaurus natans
Ophthalmosaurus icenicus
Gengasaurus nicosiai
Nannopterygius yasykovi
Nannopterygius enthekiodon
Nannopterygius saveljeviensis
Nannopterygius borealis
Arthropterygius volgensis
Arthropterygius lundi
Arthropterygius thalassonotus
Arthropterygius hoybergeti
Arthropterygius chrisorum
Brachypterygius extremus
Aegirosaurus leptospondylus
Muiscasaurus catheti
Leninia stellans
Sveltonectes insolitus
Athabascasaurus bitumineus
Platypterygius americanus
Acuetzpalin carranzai
Platypterygius sachicarum
Caypullisaurus bonapartei
Grendelius mordax
Grendelius alekseevi
Grendelius pseudoscythicus
Grendelius zhuravlevi
Undorosaurus kielanae
Undorosaurus nessovi
Undorosaurus gorodischensis
Platypterygius australis
Plutoniosaurus bedengensis
Simbirskiasaurus birjukovi
Platypterygius hercynicus
Sisteronia seeleyi
Platypterygius platydactylus
Maiaspondylus lindoei
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