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↑ 11,011,1Schultz, L. P., 1949. A further contribution to the ichthyology of Venezuela. Proceedings of the United States National Museum v. 99 (núm. 3235): 1-211, Pls. 1-3.
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↑Fowler, H. W., 1943. A collection of fresh-water fishes from Colombia, obtained chiefly by Brother Nicéforo Maria. Proc. Acad. Nat. Sci. Phila. v. 95: 223-266.
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