In 2007, a phylogenetic study of DNA sequences showed that Pleocnemia should be transferred from the Tectariaceae to the Dryopteridaceae.[9] In 2010, in a paper on bolbitidoid ferns, Arthrobotrya was resurrected from Teratophyllum.[10] Later that year, Mickelia was described as a new genus.[11]
Some species have been removed from the genus Oenotrichia because they do not belong there or even in the family Dennstaedtiaceae where Oenotrichiasensu stricto is placed. These species probably belong in the Dryopteridaceae, but have not yet been given a generic name.[7]
In 2012, a phylogenetic study of Dryopteris and its relatives included Acrophorus, Acrorumohra, Diacalpe, Dryopsis, Nothoperanema, and Peranema within that genus.[12] The Flora of China treatment of the family, published in 2013, used phylogenetic results to sink Lithostegia and Phanerophlebiopsis into Arachniodes.[13]
The Dryopteridaceae Herter, under the classification system of Christenhusz and Chase (2014), were submerged as subfamily Dryopteridoideae Link, one of eight subfamilies constituting family Polypodiaceae. This family corresponds to the clade eupolypods I.[14] The Pteridophyte Phylogeny Group classification of 2016 (PPG I) retained the family.[1]
Phylogeny
The following cladogram for the suborder Polypodiineae (eupolypods I), based on the consensus cladogram in the Pteridophyte Phylogeny Group classification of 2016 (PPG I),[1] shows a likely phylogenetic relationship between Dryopteridaceae and the other families of the clade.
^Sue Olsen. 2007. Encyclopedia of Garden Ferns Timber Press: Portland, OR, USA. ISBN978-0-88192-819-8
^ abcSmith et al., 2006Archived February 26, 2008, at the Wayback Machine Alan R. Smith, Kathleen M. Pryer, Eric Schuettpelz, Petra Korall, Harald Schneider & Paul G. Wolf: "A classification for extant ferns," Taxon, 55(3): 705–731 (Aug 2006)
^Eric Schuettpelz and Kathleen M. Pryer. 2009. "Evidence for a Cenozoic radiation of ferns in an angiosperm-dominated canopy". Proceedings of the National Academy of Sciences 106(27):11200-11205. doi:10.1073/pnas.0811136106
^Karl U. Kramer (with Richard E. Holttum, Robin C. Moran, and Alan R. Smith). 1990. "Dryopteridaceae". pages ??. In: Klaus Kubitzki (general editor); Karl U. Kramer and Peter S. Green (volume editors) The Families and Genera of Vascular Plants volume I. Springer-Verlag: Berlin;Heidelberg, Germany. ISBN978-0-387-51794-0
^ abAlan R. Smith, Kathleen M. Pryer, Eric Schuettpelz, Petra Korall, Harald Schneider, and Paul G. Wolf. 2008. "Dryopteridaceae". pages ??. In: "Fern Classification". pages 417-467. In: Tom A. Ranker and Christopher H. Haufler (editors). Biology and Evolution of Ferns and Lycophytes. Cambridge University Press. ISBN978-0-521-87411-3
^Hong-Mei Liu, Xian-Chun Zhang, Wei Wang, Yin-Long Qiu, and Zhi-Duan Chen. 2007. "Molecular Phylogeny of the Fern Family Dryopteridaceae inferred from Chloroplast rbcL and atpB genes". International Journal of Plant Sciences168(9):1311-1323. doi:10.1086/521710
^Robbin C. Moran, Paulo H. Labiak, and Michael Sundue. 2010. "Phylogeny and character evolution of the bolbitidoid ferns (Dryopteridaceae)". International Journal of Plant Sciences171(5):547-559. doi:10.1086/652191
^Robbin C. Moran, Paulo H. Labiak, and Michael Sundue. 2010. "Synopsis of Mickelia, a newly recognized genus of bolbitidoid ferns (Dryopteridaceae)". Brittonia62(4):337-356.
^Li-Bing Zhang, Liang Zhang, Shi-Yong Dong, and Atsushi Ebihara. 2012. "Molecular circumscription and major evolutionary lineages of the fern genus Dryopteris (Dryopteridaceae)". BMC Evolutionary Biology12(1):180
^He H, Wu SG, Xiang JY, Barrington DS (2013) "Arachniodes". In: Wu ZY, Raven PH, Hong DY (eds) Flora of China, vol 2–3.