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On 1 September 1933, he was appointed executive officer of the cruiser Iwate, and on 16 November 1936—after his promotion to captain—he was given his first command: the aircraft carrierHōshō. In 1939, he became captain of the aircraft carrier Akagi.
Promoted to rear admiral on 15 November 1940, he was commander of the 24th Air Flotilla before being appointed Chief of Staff of the 1st Air Fleet under Admiral Chūichi Nagumo in April 1941. He was involved in strategic and tactical planning and execution, including the attack on Pearl Harbor and Battle of Midway. Kusaka badly sprained both ankles and was burned during the evacuation from the critically damaged Akagi during the battle. During the aftermath of the battle, Kusaka was able to dissuade the fleet commander and senior officers from committing suicide after the Japanese defeat.[2]
Kusaka remained with the fleet until November 1942, and accepted a number of staff positions thereafter. He was promoted to vice admiral on 1 May 1944 and transferred to the Combined Fleet as Chief of Staff under commander-in-chief Admiral Soemu Toyoda in November 1944. His final assignment was command of the 5th Air Fleet after the suicide of Matome Ugaki, exactly the day Japan surrendered to the Allies.
Kusaka was interviewed by Walter Lord and gave detailed accounts for the books Day of Infamy (1957) and Incredible Victory (1967).