Nemegt Formation

Nemegt Formation
Stratigraphic range: Maastrichtian
~70 Ma
↓
Nemegt, the type locality of the Nemegt Formation
TypeGeological formation
Sub-unitsLower, Middle and Upper units
UnderliesAlluvium
OverliesBarun Goyot Formation
ThicknessOver 235 m (771 ft)
Lithology
PrimaryShale, sandstone
OtherMudstone, conglomerate
Location
Coordinates43°30′N 101°00′E / 43.5°N 101.0°E / 43.5; 101.0
Approximate paleocoordinates40°48′N 90°12′E / 40.8°N 90.2°E / 40.8; 90.2
RegionBayankhongor Aimag, Omnogov, Ovorkhangai
Country Mongolia
ExtentGobi Desert

Cretaceous-aged dinosaur fossil localities of Mongolia. Nemegt localities at area A.

The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians, and a diverse fauna of dinosaurs, including birds.

Description

The Nemegt Formation is composed of mudstones and sandstones that were deposited by ancient lakes, streams, and flood plains. The Altan Uul locality was described by Michael Novacek as "a canyon carved out of a very rich series of sedimentary rocks" with "steep cliffs and narrow washes". The climate associated with it was wetter than when preceding formations were deposited; there seems to have existed at least some degree of forest cover. Fossilized trunks have been also found. These petrified wood, and the remains of Araucariaceae conifers indicate that the forests of the Nemegt were thickly wooded, with a high canopy formed by tall conifer trees. When examined, the rock facies of this formation suggest the presence of stream and river channels, mudflats, and shallow lakes. Sediments also indicate that there existed a rich habitat, offering diverse food in abundant amounts that could sustain massive Cretaceous dinosaurs.[1][2]

Stratigraphy

Nemegt and Barun Goyot formations intersection

The most recent stratigraphy divides the Nemegt Formation into three informal members. The lower member is dominated by fluvial deposits, while middle and upper members consist of alluvial plain, paludal, lacustrine, and fluvial deposits. The colour of the sediments is usually light grey to tan in colour in comparison to the typically red colour of the underlying Barun Goyot Formation.[3] It overlies and sometimes interfingers with the Barun Goyot Formation. Interfingering has been noted at the stratotype (Red Walls) and Hermiin Tsav.[3] There has been no absolute dating of the Nemegt Formation. Historically the Nemegt has been considered late Campanian to Maastrichtian, based on comparisons of fossils present, but no exact dating has been performed. The age for the underlying Baruungoyot Formation (= Svita) has been suggested as Santonian to Campanian, and Shuvalov (2000) found K-Ar dating of basalts that they referred to the mid and upper Baruungoyot to be 75 to 80 million years old.[4]

Paleobiota of the Nemegt Formation

Stratigraphic positions are based on Eberth (2018) who correlated localities to their approximate position within the formation.[3]

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Amphibians

Genus Species Location Stratigraphic position Material Notes Image
Altanulia A. alifanovi Altan Uul II Middle or Upper "Partial maxilla."[5] A discoglossid. Its classification has been criticized.[6]

Crocodylomorphs

Genus Species Location Stratigraphic position Material Notes Image
Paralligator[7] P. gradilifrons[7] Nemegt, Nogon Tsav, Ulaan Bulag Lower "Partially complete skulls with sparse body elements."[7] A paralligatorid.

Fish

Genus Species Location Stratigraphic position Material Notes Image
Harenaichthys H. lui Altan Uul, Bügiin Tsav, Hermiin Tsav, Ulan Khushu "Partial skull parts, isolated and articulated centra, and articulated caudal fin."[8] An osteoglossomorph. Includes previously reported material.[9][8]
Hiodontidae indet. Indeterminate Ulan Khushu "Partial vertebrae and premaxilla."[9] A mooneye.
Teleostei indet. Indeterminate Not specified. "Vertebral centra found within a specimen of Raptorex."[9] A teleost.
Osteichthyes indet. Indeterminate Bügiin Tsav "Vertebrae and scales found within a specimen of Deinocheirus''."[10] A bony fish.

Flora

Genus Species Location Stratigraphic position Material Notes Image
Araucariaceae indet. Indeterminate Altan Uul II, Nemegt "Petrified wood and trunks."[11][2] An araucarian.
Monilitheca M. minuta Ulaan Bulag "Two isolated megasporophylls and megaspore casts."[12] A sporophyll related to quillworts.
Lemnaceae indet. Indeterminate Tsagan Khushu, Ulaan Bulag "Fruits."[13] Duckweed.
Potamogeton-like indet. Indeterminate Ulaan Bulag "Shoots and leaves."[13] Pondweed.

Invertebrates

Genus Species Location Stratigraphic position Material Notes Image
Altanicypris A. bispinifera Altan Uul IV, Nemegt Lower "Complete carapace and left valve."[14] An ostracod.
A. multispina Altan Uul IV, Nemegt Lower "Nine carapaces and some detached valves."[14] An ostracod.
A. szczechurae Altan Uul IV, Nemegt Lower "Multiple carapaces and detached valves."[14] An ostracod.
Candona C. altanulaensis Altan Uul IV, Bügiin Tsav, Nemegt "Multiple specimens with carapaces and valves."[14] An ostracod.
cf. C. fabaeformis Altan Uul IV, Nemegt Lower "Several carapaces and detached valves."[14] An ostracod.
Candoniella C. altanica Altan Uul IV, Bügiin Tsav, Nemegt "Multiple specimens with carapaces and valves."[14] An ostracod.
C. mordvilkoi Altan Uul IV Lower "Three adult carapaces."[15] An ostracod.
Cyclocypris C. transitoria Bügiin Tsav, Nemegt "Seven adult carapaces."[14] An ostracod.
Cypria C. elata Bügiin Tsav, Hermiin Tsav "Several carapaces and valves."[14] An ostracod.
Cypridopsis? C. bugintsavicus Altan Uul IV, Bügiin Tsav, Nemegt, Tsagan Khushu "Ten adult carapaces."[14] An ostracod.
Indeterminate Nemegt Lower "Several complete carapaces."[14] An ostracod.
Cypris? C. ectypa Altan Uul IV, Bügiin Tsav, Nemegt, Ulan Bulag "Complete carapaces and some detached valves."[14] An ostracod.
Cypridea cf. C. punctilataeformis Nemegt "One adult carapace."[15] An ostracod.
C. barsboldi Altan Uul IV, Bügiin Tsav, Nemegt "Multiple specimens with carapaces and valves."[14] An ostracod.
C. cavernosa Altan Uul IV, Bügiin Tsav, Nemegt, Nogon Tsav "Multiple specimens with carapaces and valves."[14] An ostracod.
C. rostrata Bügiin Tsav, Nemegt, Nogon Tsav "Caparaces."[15] An ostracod.
Indeterminate Altan Uul IV, Nemegt Lower "Five juvenile carapaces."[14] An ostracod.
Gobiella G. prima Altan Uul IV, Nemegt Lower "Several carapaces and detached valves."[14] An ostracod.
Khandia K. stankevitchae Altan Uul IV, Nemegt Lower "Multiple carapaces and valves."[14] An ostracod.
Leiria Indeterminate Nemegt Lower "Several detached valves."[14] An ostracod.
Indeterminate Nemegt Lower "Complete carapace."[14] An ostracod.
Limnocythere sp. Indeterminate Nemegt Lower "Juvenile carapace."[14] An ostracod.
Lycopterocypris? cf. L. profunda Altan Uul IV, Nemegt Lower "Multiple complete carapaces and left valve."[14] An ostracod.
Mongolianella M. palmosa? Altan Uul IV, Bügiin Tsav, Nemegt "Several adult carapaces."[14] An ostracod.
Mongolocypris M. distributa Altan Uul IV, Bügiin Tsav, Nemegt, Tsagan Khushu "Multiple specimens with complete carapaces and valves."[14] An ostracod.
Nemegtia N. biformata Altan Uul IV, Bügiin Tsav, Nemegt, Tsagan Khushu "Multiple complete carapaces and several detached valves."[14] An ostracod.
N. obliquecostae Altan Uul IV, Nemegt Lower "Multiple carapaces and valves."[14] An ostracod.
N. reticulata Altan Uul IV, Bügiin Tsav, Nemegt, Nogon Tsav, Tsagan Khushu "Several carapaces and detached valves."[14] An ostracod.
Paracypridea? P. mongolica Altan Uul IV, Nemegt Lower "Multiple carapaces and detached valves."[14] An ostracod.
Rhinocypris sp. Indeterminate Nemegt Lower "Several carapaces."[14] An ostracod.
Scabriculocypris S. ingenicus? Altan Uul IV, Bügiin Tsav, Naran Bulag, Nemegt, Tsagan Khushu "Multiple carapaces and valves."[14] An ostracod.
S. rasilis? Altan Uul IV, Nemegt Lower "Several carapaces and detached valves."[14] An ostracod.
Timiriasevia cf. T. miaogouensis Altan Uul IV, Nemegt Lower "Seven valves."[14] An ostracod.
cf. T. opinabilis Altan Uul IV Lower "Adult carapace."[15] An ostracod.
T. minuscula Altan Uul IV, Bügiin Tsav, Nemegt "Multiple specimens with carapaces and valves."[14] An ostracod.
T. naranbulakensis Altan Uul IV, Bügiin Tsav, Nemegt "Four carapaces and one right valve."[14] An ostracod.
Ziziphocypris Z. costata Altan Uul IV, Bügiin Tsav, Nemegt "Fourteen carapaces."[14] An ostracod.

Mammals

Genus Species Location Stratigraphic position Material Notes Image
Buginbaatar[16] B. transaltaiensis[16] Bügiin Tsav Middle or Upper "Fragmentary skull."[16] A multituberculate.
"Gurlin Tsav skull" Indeterminate Middle or Partial skull A mysterious metatherian, possibly a sparassodont.

Pterosaurs

Genus Species Location Stratigraphic position Material Notes Image
Unnamed azhdarchid Indeterminate Guriliin Tsav Middle or Upper "Fragmentary cervical vertebrae."[17] A giant azhdarchid.

Turtles

Genus Species Location Stratigraphic position Material Notes Image
Gobiapalone G. breviplastra Alak Shand Khuduk, Altan Uul I, Bügiin Tsav, Bügiin Tsav II, Nemegt, Ulan Khushu "Shell and body elements from multiple specimens."[18] A trionychid.
Gravemys G. barsboldi Bamba Khuduk, Hermiin Tsav, Ingeni Khobur, Tsagan Khushu "Partial to complete shells."[19][20] A lindholmemydid.

Nanhsiungchelyidae indet.

Indeterminate Hermiin Tsav II Lower "Shell fragment."[21] A nanhsiungchelyid.
Indeterminate Khuren Tsav "Damaged shell."[21] A nanhsiungchelyid.
Indeterminate Nemegt Lower "Partial plastron."[21] A nanhsiungchelyid.
Mongolemys M. elegans Bügiin Tsav, Hermiin Tsav, Tsagan Khushu "Skull, shell and body elements from multiple specimens."[22][23] A lindholmemydid.
Mongolochelys M. efremovi Altan Uul II, Altan Uul III, Bügiin Tsav, Guriliin Tsav, Hermiin Tsav, Nemegt, Tsagan Khushu "Skull, shell and body elements from multiple specimens."[24][25] A sichuanchelyid.
Nemegtemys N. conflata Bügiin Tsav, Nemegt "Partial plastron elements."[18] A trionychid.

"Trionyx"

"T". gilbentuensis Gilbentu "Fragmentary shell."[18] A trionychid.
"T". gobiensis Altan Uul III, Bamba Khuduk, Nemegt, Tsagan Khushu "Fragmentary shell and carapace fragments."[18] A trionychid.
Trionychidae indet. Indeterminate Bamba Khuduk, Bügiin Tsav, Guriliin Tsav, Ingeni Khobur "Shell and body remains."[18] A trionychid. Originally identified as Amyda menneri.[18]

Dinosaurs

Ornithischians

Ankylosaurs
Genus Species Location Stratigraphic position Material Notes Image
"Dyoplosaurus"[26] "D". giganteus[26] Nemegt Lower A series of caudal vertebrae, metatarsals, phalanges, osteoderms, and an undescribed partial tail club.[27] An ankylosaurid now regarded as nomen dubium.
Saichania[28] S. chulsanensis[28] Altan Uul IV Lower Partial vertebrae, tail club, and osteoderms.[29] An ankylosaurid also present in the Barun Goyot Formation.
Tarchia[28] T. teresae[29] Altan Uul IV, Hermiin Tsav I Lower A skull, mandibles, vertebrae, tail club, and other undescribed postcranial material.[29] An ankylosaurid.
T. tumanovae[30] Hermiin Tsav Lower[3] "Partial skeleton with complete skull."[30] An ankylosaurid.
Tetrapodosaurus-like indet. Indeterminate Bügiin Tsav, Shar Tsav "Footprint casts."[31] Ankylosaurid tracks.
Ankylosauridae indet. Indeterminate Altan Uul II-III-IV, Bügiin Tsav, Hermiin Tsav I, Khuree Tsav, Nemegt A partial dentary, caudal vertebrae, dorsal vertebrae, metatarsals, phalanges, pedal phalanx, humeri, partial pelvis, tail clubs, cervical half-rings, osteoderms, skin impressions, and undescribed juveniles and partial skeletons.[27] Cranial and postcranial material representing numerous individuals of different ages.
Hadrosaurs
Genus Species Location Stratigraphic position Material Notes Image
Amblydactylus-like indet. Indeterminate Bügiin Tsav, Bügiin Tsav II, Guriliin Tsav, Yagaan Khovil "Footprint casts."[31] Three-toed hadrosaurs tracks. Attributed to Saurolophus.
Barsboldia[32] B. sicinskii[32] Nemegt Lower[3] "Partial vertebrae, pelvis, and ribs."[32][33] A hadrosaurid.
Hadropodus Indeterminate Nemegt "Numerous footprints."[34] Hadrosaurid tracks. Attributed to Saurolophus.
Saurolophus[35] S. angustirostris[36] Presence in most localities[37] "Multiple specimens, including articulated skulls, postcranial skeletons and mass death assemblage."[38][39][40][41] A saurolophine hadrosaurid which was the most common hadrosaur in the area.[42]
Unnamed ichnotaxon Indeterminate Nemegt "Numerous footprint casts with trackways."[43] Hand and feet hadrosaur tracks. Attributed to Saurolophus.
Pachycephalosaurs
Genus Species Location Stratigraphic position Material Notes Image
Homalocephale[44] H. calathocercos[44] Nemegt Lower[3] "Partial skull and skeleton including hindlimbs."[44] A pachycephalosaurid.
Prenocephale[44] P. prenes[44] Guriliin Tsav, Nemegt "Complete skull and partial postcranium of several specimens."[44][46] A pachycephalosaurid.
P. sp.[46] Tsagan Khushu, Nemegt Lower[3] "complete left squamosal"; large skull
Indeterminate Bugiin Tsav Middle[45] "incomplete frontoparietal dome and fragments of associated braincase bones"[46]

Sauropods

Genus Species Location Stratigraphic position Material Notes Image
Brontopodus Indeterminate Nemegt Lower "Large footprint and several four-toed tracks."[47][34] Sauropod tracks. Attributed to Opisthocoelicaudia.
Brontopodus-like indet. Indeterminate Bügiin Tsav, Yagaan Khovil "Footprint casts."[31] Sauropod tracks.
Nemegtosaurus[48] N. mongoliensis[48] Nemegt Lower "Skulls and body remains from several specimens."[49][50][51] A titanosaur.
Opisthocoelicaudia[52] O. skarzynskii[52] Altan Uul IV Lower "Nearly complete skeleton without the skull and cervical series."[50] A titanosaur.
Undescribed sauropods Indeterminate Altan Uul I-II-II-IV, Bügiin Tsav, Guriliin Tsav, Hermiin Tsav, Khamaryn Khural, Khuree Tsav, Nemegt, Nogon Tsav, Tsagan Khushu, Ulan Khushu "Several fragmentary remains."[50] Sauropod remains that may represent already named taxa.
Unnamed ichnotaxon Indeterminate Nemegt "Footprint casts with skin impressions."[43] Four-toed sauropod tracks. Attributed to Opisthocoelicaudia.

Theropods

Alvarezsaurs
Genus Species Location Stratigraphic position Material Notes Image
Mononykus[53] M. olecranus[53] Bügiin Tsav Middle[45] "Fragmentary skull, vertebrae, limbs and fragmented pelvis."[54] An alvarezsaurid.
Indeterminate Altan Uul III "Caudal vertebrae and left hindlimb."[55]
Nemegtonykus[55] N. citus[55] Altan Uul III "Partially complete skeleton from two specimens lacking the skull."[55] An alvarezsaurid.
Birds
Genus Species Location Stratigraphic position Material Notes Image
Brodavis[56] B. mongoliensis[56] Bügiin Tsav "Nearly complete left tarsometatarsus."[56] A hesperornithine.
Gurilynia[57] G. nessovi[57] Guriliin Tsav Middle[45] "Left coracoid and partial humerii."[57] An enantiornithine.
Judinornis[58] J. nogontsavensis[58] Nogon Tsav "Dorsal vertebra."[58] A hesperornithine.
Laevisoolithus[59] L. sochavai "Whole egg with partly broken pole"[59] Laid by a bird or small theropod.[59]
Subtiliolithus[59] S. microtuberculatus[59] "Eggshell fragments"[59]
Teviornis[60] T. gobiensis[60] Guriliin Tsav Middle[45] "Partial forelimbs."[60] A presbyornithid anseriform.
Unnamed ornithurans Indeterminate Tsagan Khushu "A humerus and two tibiotarsi."[61] Ornithuran remains.
Dromaeosaurs
Genus Species Location Stratigraphic position Material Notes Image
Adasaurus[62] A. mongoliensis[62] Bügiin Tsav Middle[45] "Partial skull and fragmentary postcrania from two specimens."[63] A dromaeosaurid.

Dromaeosauridae indet.

Indeterminate Bügiin Tsav Middle[45] "Cervical vertebra."[64] A dromaeosaurid.
Indeterminate Hermiin Tsav Lower "Dorsal vertebra."[65]
Indeterminate Khaichin I "Three teeth."[66]
Indeterminate Nemegt Lower "Dorsal vertebra and other elements."[65][67]
Indeterminate Indeterminate Bügiin Tsav "Footprints with poor claw marks."[68] Two-toed theropod tracks. Attributed to either Adasaurus or Zanabazar.
Ornithomimosaurs
Genus Species Location Stratigraphic position Material Notes Image
Anserimimus[69] A. planinychus[69] Bügiin Tsav Middle or Upper[3] "Partial skeleton lacking the skull."[69][70] An ornithomimid.
Deinocheirus[71] D. mirificus[71] Altan Uul III-IV, Bügiin Tsav "Complete skull with virtually complete postcranial remains from three specimens."[10] A giant deinocheirid.
Gallimimus[72] G. bullatus[72] Presence in most localities[73] "Multiple specimens with nearly complete skull and postcranial elements."[73][74] An ornithomimid.
Indeterminate Indeterminate Bügiin Tsav "Nine trackways including an articulated Gallimimus foot."[68] Three-toed theropod tracks. Attributed to ornithomimids.
Unnamed ornithomimid[75] Indeterminate Tsagan Khushu "Partial vertebral column with fore and hindlimbs."[75][73] An ornithomimid.
Oviraptorosaurs
Genus Species Location Stratigraphic position Material Notes Image
Avimimus[76] A. portentosus[76] Shar Tsav "Skull roof with fragmentary body elements."[77] An avimimid. Also present in the Djadokhta Formation.
A. nemegtensis[77] Nemegt Lower[3] "Fragmented skull with partial skeleton."[77] An avimimid.
Conchoraptor C. gracilis Guriliin Tsav Middle[45] "Skull and a flock of articulated individuals."[78] An oviraptorid. Also present in the Barun Goyot Formation.
Elmisaurus[79] E. rarus[79] Altan Uul II, Nemegt
  • Lower
  • Middle or Upper
 "Fragmentary skull and partially skeletons from several specimens."[77][80] A caenagnathid.
Elongatoolithidae indet. Indeterminate Bügiin Tsav "Three eggs containing embryonic remains."[81] Oviraptorid eggs.
Gobiraptor[82] G. minutus[82] Altan Uul III "Partial crania with complete lower jaws and fragmented postcrania."[82] An oviraptorid.
Indeterminate Indeterminate Hermiin Tsav, Shar Tsav, Yagaan Khovil "Footprint casts."[31] Three-toed theropod tracks. Attributed to Avimimus.
Nemegtomaia[83] N. barsboldi[83] Nemegt Lower "Multiple specimens including a nesting specimen and eggs."[84] An oviraptorid also present in the Barun Goyot Formation.
Nomingia[85] N. gobiensis[85] Bügiin Tsav, Nemegt "Vertebral series with partial hindlimbs and a referred pygostyle."[77] A caenagnathid. May be synonymous with Elmisaurus.[80]
Oksoko[86] O. avarsan Bügiin Tsav, Guriliin Tsav Middle[45] "Multiple associated skeletons."[86] An oviraptorid.
Rinchenia[87] R. mongoliensis[87] Altan Uul II Middle or Upper "Skull with nearly complete skeleton."[77] An oviraptorid.


Therizinosaurs
Genus Species Location Stratigraphic position Material Notes Image
Therizinosaurus[88] T. cheloniformis[88] Altan Uul, Hermiin Tsav, Nemegt Lower[3] "Fore and hindlimb elements from multiple specimens."[89][90] A giant therizinosaurid.
Undescribed therizinosaurids Indeterminate Altan Uul, Bügiin Tsav, Guriliin Tsav, Ulan Khushu "Partial remains."[91] Therizinosaurid remains.
Troodontids
Genus Species Location Stratigraphic position Material Notes Image
Borogovia[92] B. gracilicrus[92] Altan Uul IV Lower "Partial hindlimbs."[92] A troodontid.
Indeterminate Indeterminate Bügiin Tsav "Footprints with poor claw marks."[68] Two-toed theropod tracks. Attributed to either Adasaurus or Zanabazar.
Tochisaurus[93] T. nemegtensis[93] Nemegt Lower "Left metatarsus."[93] A troodontid.
Zanabazar[94] Z. junior[95] Bügiin Tsav Middle[45] "Skull with fragmentary postcranial skeleton."[94] A troodontid originally identified as "Saurornithoides junior".
Tyrannosaurs
Genus Species Location Stratigraphic position Material Notes Image
Alioramus[96] A. remotus[96] Nogon Tsav "Partially preserved skull with very sparse postcranial remains."[96] An alioramin.
A. altai[97] Tsagan Khushu "Nearly complete skull with partial postcrania."[97] An alioramin.
Bagaraatan[98] B. ostromi[98] Nemegt Lower[3] "Fragmentary lower jaw, hindlimb and caudal vertebrae."[98] A tyrannosauroid.
Indeterminate Indeterminate Bügiin Tsav, Bügiin Tsav II "Footprint casts."[31] Three-toed theropod tracks. Attributed to Tarbosaurus.

Raptorex[99]

R. kriegsteni[99]

"Nearly complete skeleton including the skull."[99] A controversial genus of tyrannosaurid that could be a juvenile Tarbosaurus.[100]
Tarbosaurus[101] T. bataar[102] Presence in most localities[2] "Nearly complete skeletons from multiple specimens."[101][102][2] A large tyrannosaurid which was the most common large carnivore in the area.[103]
Unnamed ichnotaxon Indeterminate Nemegt "Footprint casts with skin impressions."[43] Three-toed theropod tracks. Attributed to Tarbosaurus.
Tyrannosauripodidae indet. Indeterminate Nemegt "Five tracks."[34] Three-toed tyrannosaurid tracks. Attributed to either Alioramus or Tarbosaurus.

See also

References

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