Macrobaenidae

Macrobaenidae; Temporal range: Early Cretaceous–Paleocene PreꞒ Ꞓ O S D C P T J K Pg N
	Osteopygis emarginatus carapace
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Clade:	Pantestudines
Clade:	Testudinata
Clade:	Perichelydia
Family:	†Macrobaenidae; Sukhanov, 1964
Genera
	See text

Macrobaenidae is an extinct family of turtles, known from the Early Cretaceous to Paleogene of Laurasia. Their relationships to other turtles and whether they form a monophlyletic group are controversial. They are typically interpreted as stem or crown group cryptodires, but some more recent analyses have found them to lie outside crown group Testudines. Macrobaenids can be distinguished from other testudinatans by the presence of a carotid fenestra, cruciform plastron with strap-like epiplastra, and a lack of extragulars.^[1]

Genera

Anatolemys^[2] Central Asia, Late Cretaceous ( Khodzhakul Formation, Uzbekistan, Cenomanian, Bissekty Formation, Uzbekistan, Turonian, Bostobe Formation, Kazakhstan, Santonian Yalovach Formation, Tajikistan, Santonian)
Appalachemys^[3] Mooreville Chalk, Alabama, Late Cretaceous (Santonian–Campanian)
Asiachelys^[2] Khulsangol Formation, Mongolia, Early Cretaceous (Albian)
Aurorachelys^[2] Kanguk Formation, Canada,Turonian
Changmachelys Xiagou Formation, China, Early Cretaceous (Aptian)
Kirgizemys^[2] Murtoi Formation, Khilok Formation, Ilek Formation, Russia, Early Cretaceous (Barremian-Aptian), Alamyshik Formation, Kyrgyzstan, Albian, Geoncheonri Formation, South Korea, Albian Khodzhakul Formation, Uzbekistan, Cenomanian
Judithemys North America, Late Cretaceous–Paleocene
Gallica^[4] Paris Basin, France, Paleocene
Macrobaena^[2] Naran Bulak Formation, Mongolia, Paleocene
Osteopygis^[2] Late Cretaceous-Paleocene, North America
Oxemys^[2] Khodzhakul Formation, Uzbekistan, Cenomanian
Yakemys Phu Kradung Formation, Thailand, Early Cretaceous (Berriasian)

Hongkongochelys from the Middle-Late Jurassic of China has sometimes been attributed to the family,^[4] but other times has been attriuted to Sinemydidae, a group which has an unresolved relationship with Macrobaenidae.

References

^ Joyce, Walter G.; Anquetin, Jérémy; Cadena, Edwin-Alberto; Claude, Julien; Danilov, Igor G.; Evers, Serjoscha W.; Ferreira, Gabriel S.; Gentry, Andrew D.; Georgalis, Georgios L.; Lyson, Tyler R.; Pérez-García, Adán (December 2021). "A nomenclature for fossil and living turtles using phylogenetically defined clade names". Swiss Journal of Palaeontology. 140 (1): 5. doi:10.1186/s13358-020-00211-x. hdl:11336/155192. ISSN 1664-2376.
^ ^a ^b ^c ^d ^e ^f ^g "Macrobaenidae". FossilWorks. Retrieved 17 December 2021.
^ Gentry, A. D.; Kiernan, C. R.; Parham, J. F. (2022). "A large non-marine turtle from the Upper Cretaceous of Alabama and a review of North American "Macrobaenids"". The Anatomical Record. doi:10.1002/ar.25054.
^ ^a ^b Adán Pérez-García (2020). "A European Cenozoic 'Macrobaenid:' New Data about the Paleocene Arrival of Several Turtle Lineages to Europe". Journal of Vertebrate Paleontology. 40 (4): e1795874. doi:10.1080/02724634.2020.1795874. S2CID 225151817.