The Late Pleistocene is an unofficial age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a stratigraphic perspective. It is intended to be the fourth division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently defined as the time between c. 129,000 and c. 11,700 years ago. The late Pleistocene equates to the proposed Tarantian Age of the geologic time scale, preceded by the officially ratified Chibanian (commonly known as the Middle Pleistocene).[4] The beginning of the Late Pleistocene is the transition between the end of the Penultimate Glacial Period and the beginning of the Last Interglacial around 130,000 years ago (corresponding with the beginning of Marine Isotope Stage 5).[5] The Late Pleistocene ends with the termination of the Younger Dryas, some 11,700 years ago when the Holocene Epoch began.[6]
The term Upper Pleistocene is currently in use as a provisional or "quasi-formal" designation by the International Union of Geological Sciences (IUGS). Although the three oldest ages of the Pleistocene (the Gelasian, the Calabrian and the Chibanian) have been officially defined, the late Pleistocene has yet to be formally defined.[7]
Following the brief Last Interglacial warm period (~130–115,000 years ago), where temperatures were comparable to or warmer than the Holocene, the Late Pleistocene was dominated by the cool Last Glacial Period, with temperatures gradually lowering throughout the period, reaching their lowest during the Last Glacial Maximum around 26–20,000 years ago.
The Last Glacial Maximum was reached during the later millennia of the Würm/Weichselian, estimated between 26 ka and 19 ka when deglaciation began in the Northern Hemisphere. The Würm/Weichselian endured until 16 ka with Northern Europe, including most of Great Britain, covered by an ice sheet. The glaciers reached the Great Lakes in North America.[6] Sea levels fell and two land bridges were temporarily in existence that had significance for human migration: Doggerland, which connected Great Britain to mainland Europe; and the Bering land bridge which joined Alaska to Siberia.[9][10]
The end of the Younger Dryas marks the boundary between the Pleistocene and Holocene Epochs. Hominids in all parts of the world were still culturally and technologically in the Palaeolithic (Old Stone) Age. Tools and weapons were basic stone or wooden implements. Nomadic tribes followed moving herds. Non-nomadics acquired their food by gathering and hunting.[12]
Africa
Its present physical geography and climate have changed over time caused by the movement of tectonic plates and volcanoes but glacial cycles and sea level variation have a more significant effect on the vertebrate communities during the Late Pleistocene.[13]
The Late Pleistocene was the time when most animals evolved to resemble modern-day animals and they managed to live through the Late mid-Pleistocene since there were no extinction events of megafauna until the end of the Late Pleistocene.[13]
Some species which went extinct at the end of the Late Pleistocene in Southern Africa are the giant warthog, long-horn buffalo, Southern springbok, etc.[14] These species were common because their distribution changed in response to climatic influences on vegetation. Carnivores were more widespread due to their varying habitat requirements.
In Egypt, the Late (or Upper) Palaeolithic began sometime after 30,000 BC. People in North Africa had relocated to the Nile Valley as the Sahara was transformed from grassland to desert.[15] The Nazlet Khater skeleton was found in 1980 and has been radiocarbon dated to between 30,360 and 35,100 years ago.[16][17]
Most of the knowledge of the Late Pleistocene is obtained from regions like Morocco, Algeria, Tunisia, some coastal regions of Maghreb, Libya and Egypt. The only issue with interpreting the data from this region is due to the lack of chronological information.[13] The resemblance of Late Pleistocene species in Northern Africa to modern animals is the same as in Southern Africa but it's extremely difficult to date when these fauna came into place because of the lack of reliable samples from the mid-Pleistocene.[18] Most of the significant fossil records are from the Maghreb because of its geology which helps to create deep caves which is conducive for preserving fossils.
Eurasia
Neanderthal hominins (Homo neanderthalensis) inhabited Eurasia until becoming extinct between 40 and 30 ka, towards the end of the Pleistocene and possibly into the early Holocene[12][19]and were replaced with modern humans (Homo sapiens) who emerged from East Africa about 195,000 years ago.[20] Neanderthals co-existed with the Homo sapiens until they died out.[citation needed]
In Eurasia, extinction happened throughout the Pleistocene but those that happened during the Later Pleistocene were of megafauna and there were no replacements for the extinct species.[citation needed] Some Molluscan species went extinct but not on the same scale as the mammals living during the time.[21] Some examples of species which extinct without replacements include the Straight-tusked elephant (Palaeoloxodon antiquus), Giant deer (Megaloceros giganteus), cave bear (Ursus spelaeus) and woolly rhinoceros (Coelodonta antiquitatis).[22] Several large mammalian species including the mammoth, mastodon, and Irish elk became extinct.[23]
Upper Paleolithic people also made paintings and engravings on walls. Cave paintings have been found at Lascaux in the Dordogne which may be more than 17,000 years old. These are mainly buffalo, deer, and other animals hunted by humans. Later paintings occur in caves throughout the world, including Altamira, Spain, and in India, Australia, and the Sahara.[19][24]
Magdalenian hunter-gatherers were widespread in western Europe about 20–12.500 cal BP years ago until the end of the Pleistocene.[25] An example of this is the antler-working done by the human groups who lived in the Santimamine cave in the Magdalenian.[26] They invented the earliest known harpoons using reindeer horn.[27]
Climatic conditions during the Late Pleistocene in Eurasia were predominantly cold with glaciation events happening in northern Europe, northwest Siberia and the Alps and interglacials (temperate phase). The evidence of the changes in climatic conditions was from fragmentary sequences in formerly glaciated areas in northern Europe.[22]
The only domesticated animal in the Pleistocene was the dog, which evolved from the grey wolf into its many modern breeds. It is believed that the grey wolf became associated with hunter-gatherer tribes around 15 Ka.[28] The earliest remains of a true domestic dog have been dated to 14,200 years ago.[29] Domestication first happened in Eurasia but could have been anywhere from Western Europe to East Asia.[30] Domestication of other animals such as cattle, goats, pigs, and sheep did not begin until the Holocene when settled farming communities became established in the Near East.[28] The cat was probably not domesticated before c. 7500 BC at the earliest, again in the Near East.[31]
Japan was once linked to the Asian mainland by land bridges via Hokkaido and Sakhalin Island to the north but was unconnected at this time when the main islands of Hokkaido, Honshu, Kyushu and Shikoku were all separate entities.[37]
North America
Human migrations happened during this time with people coming in from Eurasia. From about 28 ka, there were migrations across the Bering land bridge from Siberia to Alaska. The people became the Native Americans. It is believed that the original tribes subsequently moved down to Central and South America under pressure from later migrations.[10][19]
Bison occidentalis and Bison antiquus, an extinct subspecies of the smaller present-day bison, survived the late Pleistocene period, between about 12 and 11 ka ago. Clovis people depended on these bison as their major food source. Earlier kills of camels, horses, and muskoxen found at Wally's beach were dated to 13.1–13.3 ka B.P.[41]
South America
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Over 50 genera (~ 83%) of megafauna in South and North America went extinct during the Pleistocene.[42] most mega mammals (>1000kg) and large mammals (>40kg) went extinct by the end of the Late Pleistocene.[43] During this period there was a major cooling event called the Younger Dryas and the Clovis culture of capturing game became more prominent.[44] Diverse factors such as climate change may have triggered this extinction but it's still in debate what the major factors were.[45]
The Late Pleistocene saw a change in the use of coastal resources and advancements in marine technology. The reasons for these changes have not been confirmed; various triggering mechanisms have been theorized such as climate change, the arrival of new people, or the struggle for resources.[46]
There is evidence of human habitation in mainland Australia, Indonesia, New Guinea and Tasmania from c. 45,000 BC. The finds include rock engravings, stone tools and evidence of cave habitation.[48]
In Australia, there are sites which show evidence of pollen records from the Late Pleistocene and they are mostly found in more temperate regions of the continent.[49] Some megafauna decreased in size over time, while others remained the same; however, the fossil record is limited in the exact chronologies of the extinctions.[50]
In general, various reasons have been stated to have caused the extinctions during the Late Pleistocene but the topic is still being debated.[51]
^Cohen, K. M.; Finney, S. C.; Gibbard, P. L.; Fan, J.-X. (January 2020). "International Chronostratigraphic Chart"(PDF). International Commission on Stratigraphy. Retrieved 23 February 2020.
^ abWinter, Barbara. "Bering Land Bridge". SFU Museum of Archaeology and Ethnology. Archived from the original on 28 April 2015. Retrieved 2 March 2019.
^Sutcliffe, Antony J. (1986). On the track of ice age mammals (Reprinted with amendments ed.). London: British Museum (Natural History). ISBN978-0-565-00869-7.
^Olaf Thalmann; Angela R. Perri (2018). "Palaeogenomic Inferences of Dog Domestication". In Lindqvist, C.; Rajora, O. (eds.). Palaeogenomics. Population Genomics. Springer, Cham. pp. 273–306. doi:10.1007/13836_2018_27. ISBN978-3-030-04752-8.
^David E. Machugh; et al. (2016). "Taming the Past: Ancient DNA and the Study of Animal Domestication". Annual Review of Animal Biosciences. 5: 329–351. doi:10.1146/annurev-animal-022516-022747. PMID27813680.
^A. E. Sanders, R. E. Weems & L. B. Albright III (2009). Formalization of the mid-Pleistocene "Ten Mile Hill beds" in South Carolina with evidence for placement of the Irvingtonian-Rancholabrean boundary. Museum of Northern Arizona Bulletin (64:369–375).
^D. E. Savage (1951). Late Cenozoic vertebrates of the San Francisco Bay region. University of California Publications; Bulletin of the Department of Geological Sciences (28:215–314).
^Bell, C. J. (2004). "The Blancan, Irvingtonian, and Rancholabrean mammal ages". In Woodburne, M. O. (ed.). Late Cretaceous and Cenozoic Mammals of North America: Biostratigraphy and Geochronology. New York: Columbia University Press. pp. 232–314. ISBN0-231-13040-6.
Ehlers, J., and P.L. Gibbard, 2004a, Quaternary Glaciations: Extent and Chronology 2: Part II North America. Elsevier, Amsterdam. ISBN0-444-51462-7
Ehlers, J., and P L. Gibbard, 2004b, Quaternary Glaciations: Extent and Chronology 3: Part III: South America, Asia, Africa, Australia, Antarctica. ISBN0-444-51593-3
Frison, George C., Prehistoric Human and Bison Relationships on the Plains of North America, August 2000, International Bison Conference, Edmonton, Alberta.
Gillespie, A. R., S. C. Porter, and B. F. Atwater, 2004, The Quaternary Period in the United States. Developments in Quaternary Science no. 1. Elsevier, Amsterdam. ISBN978-0-444-51471-4
Mangerud, J., J. Ehlers, and P. Gibbard, 2004, Quaternary Glaciations : Extent and Chronology 1: Part I Europe. Elsevier, Amsterdam. ISBN0-444-51462-7
Sibrava, V., Bowen, D. Q., and Richmond, G. M., 1986, Quaternary Glaciations in the Northern Hemisphere, Quaternary Science Reviews. vol. 5, pp. 1–514.
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