AS 4.1848, AS 4.1888, ATCC 23924, BCRC 13773, CBS 908.68, CCRC 13773, CGMCC 4.1848, CGMCC 4.1888, DSM 40124, ETH 28997, IFO 12878, IFO 3868, IMET 42938, ISP 5124, JCM 4646, KCC S-0646, KCCS-0646, NBRC 12878, NBRC 3868, NCIB 9718, NCIMB 9718, NRRL B-3301, NRRL-ISP 5124, OEU 709, RIA 1127, Shinobu 709
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