Coscinia cribraria

Speckled footman
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Lepidoptera
Superfamily:	Noctuoidea
Family:	Erebidae
Subfamily:	Arctiinae
Genus:	Coscinia
Species:	C. cribraria
Binomial name
Coscinia cribraria (Linnaeus, 1758)
Synonyms
List Phalaena (Bombyx) cribraria Linnaeus, 1758 Coscinia cribrumella J. Hübner, 1756 Euprepia cribraria (Linnaeus, 1758) Spiris cribraria (Linnaeus, 1758) Coscinia cribrum (Linnaeus, 1761) Emydia cribrum (Linnaeus, 1761) Phalaena cribrum Linnaeus, 1761 Coscinia cribellum Esper, 1786 Coscinia candida Cyrillo, 1787 Emydia cribrum subsp. candida (Cyrillo, 1787) Coscinia colon J. Hübner, 1792 Coscinia albeola J. Hübner, 1804 Coscinia chrysocephala J. Hübner, 1804 Coscinia coscinia Ochsenheimer, 1810 Emydia coscinia (Ochsenheimer, 1810) Coscinia cyrillii J. Hübner, 1816 Coscinia bifasciata (Rambur, 1832) Euprepia bifasciata Rambur, 1832 Coscinia punctigera Freyer, 1834 Coscinia rippertii Boisduval, 1834 Coscinia candida Herrich-Schäffer, 1846 Coscinia inquinata Rambur, 1866 Coscinia sibirica Staudinger, 1892 Coscinia infuscata Rey, 1903 Coscinia anglica Oberthür, 1911 Coscinia canigulensis Oberthür, 1911 Coscinia leucomelas Oberthür, 1911 Coscinia nevadensis Oberthür, 1911 Coscinia rondoni Oberthür, 1911 Coscinia vernetensis Oberthür, 1911 Coscinia pseudozatima Krulikowsky, 1911 Coscinia fortestrigata Schawerda, 1929 Coscinia paucisignata Schawerda, 1929 Coscinia transversata Schawerda, 1929 Coscinia pseudobifasciata Dannehl, 1929 Coscinia ribbei Draudt, 1931 Coscinia limbata Rondou, 1932 Coscinia quadrifasciata Reisser, 1933 Coscinia nigromarginata Lingonblad, 1950 Coscinia delimbata Cockayne, 1951 Coscinia flavescens Lempke, 1961 Coscinia punctata Lempke, 1961 Coscina benderi Marten, 1957

Coscinia cribraria, the speckled footman, is a species of moth belonging to the subfamily Arctiinae within the large family Erebidae. This moth has a widespread distribution in the Palearctic.

Taxonomy

Coscinia cribraria was first described as Phalaena (Bombyx) cribraria by Carl Linnaeus in his 1758 10th edition of Systema Naturae.^[1] Linnaeus's P. cribraria was designated as the type species of the genus Coscinia by George Hampson in 1901, the genus having been proposed by Jacob Hübner in 1819.^[2] This is a polytypic species with a number of described subspecies. For example in the United Kingdom there is an endemic subspecies C. c. bivittata occurs in Dorset ad the New Forest while a migrant subspecies from continental Europe C. c. arenaria occurs as a scarce migrant in eastern England.^[3] Other taxa from Iberia, such as Coscinia benderi , C. c. rippertii and C. c. ibicenca have also been shown to be subspecies of C. ccribaria, having previously been regarded as separate species.^[4] The genus Coscinia is placed within the subtribe Callimorphina within the tribe Arctiini in the subfamily Arctiinae, part of the large family Erebidae.^[5]

Description

Coscinia cribraria has a forewing which has a background colour of dirty white marked with black streaks along the wings and crossbands made up of dark dots, both of which vary in extent. The subspecies arenaria has an almost pure white forewing.^[6]The speckled footman has a wingspan of between 30 and 35 mm (1.2 and 1.4 in).^[7]

Distribution and habitat

Coscinia cribraria has a wide Palearctic distribution being found from Northwestern Africa Iberia and southern Great Britain east to China and Mongolia.^[1] In Europe, north of the Alps, the speckled footman is typically found on sandy heaths and open sandy pine forests. To the south of the Alps, it inhabits on dry mountain meadows with stony or rocky areas and around the Mediterranean this species can be found in habitats like maquis and in open forests.^[8]

Illustration from John Curtis's British Entomology Volume 5

Biology

Coscinia cribraria feeds on a wide variety of plants and caterpillars have been recorded feeding on bristle beny (Agrostis curtisii), bell heather (Erica cinerea), cross-leaved heath (Erica tetralix), ling (Calluna vulgaris) and bilberry (Vaccinium myrtillus). However, like other footman moths, the lichens in the genus Cladonia are an important component of the habitat and larval diet of the speckled footman. In southern England the eggs are laid in June, mainly on ling and bristle bent with a few females being observed to oviposit during daylight hours. The eggs are laid high up on the stems of the plants. The caterpillars overwinter as larvae^[3] and pupate in the June following their hatching, creating rather flimsy cocoons among grass and heather.^[9] At altitudes of less than 1,000 m (3,300 ft) in this species' range south of the Alps the speckled footman is bivoltine with imagos on the wing in May and June, as well as August and September.^[8]

Conservation

Coscinia cribaria is common in the southern part ofits range but in northern Europe it is vulnerable to the destrucion of the open sandy heaths it prefers.^[8] In the United Kingdom it is classified as nationally rare meaning that it has been recorded from fewer than 15, or fewer, hectads.^[9]

References

1. "Phalaena (Bombyx) cribraria Linnaeus, 1758". GBIF. Retrieved 29 April 2024.
2. "Coscinia Hübner, 1819". Plazi. Retrieved 29 April 2024.
3. Durwyn Liley; Tony Davis; Andy Page; et al. (2024). "An upturn in the fortunes for the Speckled Footman in Britain". British Wildlife. 35 (5): 319–327.
4. Macià, Ramon; Mally, Richard; Ylla, Josep; Gastón, Javier & Huertas, Manuel (2019). "Integrative revision of the Iberian species of Coscinia Hübner, [1819] sensu lato and Spiris Hübner, [1819], (Lepidoptera: Erebidae, Arctiinae)". Zootaxa. 4615 (3): 401–449. doi:10.11646/zootaxa.4615.3.1. PMID 31716329.
5. Dacosta, M.A. & Weller, S.J. (2005). "Phylogeny and classification of Callimorphini (Lepidoptera: Arctiidae: Arctiinae)". Zootaxa. 1025: 1–94. doi:10.11646/zootaxa.1025.1.1.
6. Michael Chinery (1989). David Attenborough (ed.). Butterflies and Day Flying Moths of Britain and Europe. Collins New Generation Guide. William Collins & Sons Ltd. pp. 188–189. ISBN 0-00219785-5.
7. "72.032 BF2053 Speckled Footman Coscinia cribraria (Linnaeus, 1758)". UK Moths. Retrieved 29 April 2024.
8. "Coscinia cribraria (Linnaeus, 1758)". Lepidoptera and their ecology. Wolfgang Wagner. Retrieved 29 April 2024.
9. "Speckled Footman Coscinia cribraria". dorsetMoths.co.uk. Retrieved 29 April 2024.

External links

• Moths and Butterflies of Europe and North Africa
• Fauna Europaea
• Lepiforum e.V.