The cladogram below shows one hypothesis for the evolutionary relationships among the genera of the Pteridoideae, based on a maximum likelihood analysis using six plastid markers. The authors of the study identified four major clades. The Pteris and JAPSTT clades are found worldwide; the GAPCC clade is pantropical; and the Actiniopteris+Onychium clade is restricted to the Old World.[4]
The Pteridophyte Phylogeny Group classification of 2016 (PPG I) recognized 13 genera.[5] Shortly afterwards, the genus Gastoniella was created for three species formerly placed in Anogramma which a molecular phylogenetic analysis had shown to be distinct.[4]
^Schuettpelz et al. 2007Archived 2008-08-20 at the Wayback Machine Eric Schuettpelz, Harald Schneider, Layne Huiet, Michael D. Windham, Kathleen M. Pryer: "A molecular phylogeny of the fern family Pteridaceae: Assessing overall relationships and the affinities of previously unsampled genera." Molecular Phylogenetics and Evolution44 1172–1185 (2007)
^Schuettpelz & Pryer, 2008[permanent dead link] Eric Schuettpelz & Kathleen M. Pryer: "Ch. 15. Fern phylogeny" in Biology and Evolution of Ferns and Lycophytes], ed. Tom A. Ranker and Christopher H. Haufler. Cambridge University Press (2008)
^ abZhang, Liang; Zhou, Xin-Mao; Lu, Ngan Thi & Zhang, Li-Bing (2017). "Phylogeny of the fern subfamily Pteridoideae (Pteridaceae; Pteridophyta), with the description of a new genus: Gastoniella". Molecular Phylogenetics and Evolution. 109: 59–72. doi:10.1016/j.ympev.2016.12.037.
^PPG I (2016). "A community-derived classification for extant lycophytes and ferns". Journal of Systematics and Evolution. 54 (6): 563–603. doi:10.1111/jse.12229.
