List of non-marine molluscs of Vietnam

Location of Vietnam

The non-marine molluscs of Vietnam are a part of the molluscan fauna of Vietnam (wildlife of Vietnam). A number of species of non-marine mollusks are found in the wild in Vietnam.

There are good reasons to suppose that Vietnam, with a surface of 332,000 km², a large variety of habitats, and many different limestone 'islands' that differ from each other in faunal composition,^[1] will have a rich diversity of terrestrial molluscs.^[2] Numerous non-marine mollusc species, including more than 850 species of land gastropods,^[3] have been described from the country but many others still await discovery and description.^[2]

Freshwater gastropods

Freshwater gastropods in Vietnam include:

Neritidae

• Neritina violacea^[4]

Ampullariidae

• Pila polita^[4][5]
• Pila conica^[5]
• Pila ampullacea^[5]
• Pomacea canaliculata^[4][5]
• Pomacea insularum (d'Orbigny, 1835)^[4][5]

Viviparidae

• Cipangopaludina lecythoides^[5]
• Idiopoma umblicata^[4][5]
• Sinotaia aeruginosa^[5]
• Mekongia lithophaga^[5]
• Mekongia hainesiana^[5]
• Filopaludina sumatrensis^[5]
• Angulyagra polyzonata^[6][4][5]
• Angulyagra duchieri^[5]
• Angulyagra boettgeri^[5]

Pachychilidae

• Adamietta housei^[5]
• Adamietta reevei^[5]
• Brotia annamita^[5]
• Brotia dautzenbergiana^[5]
• Brotia hoabinhensis^[5]
• Brotia swinhoei^[5]
• Sulcospira collyra^[5]
• Sulcospira dakrongensis^[5]
• Sulcospira hainanensis^[5]
• Sulcospira proteus^[5]
• Sulcospira quangtriensis^[5]
• Sulcospira tonkiniana^[5]
• Sulcospira touranensis^[5]
• Sulcospira vietnamensis^[5]
• Semisulcospira aubryana^[5]
• Paracrostoma paludiformis^[5]

Paludomidae

• Paludomus messageri (Bavay & Dautzenberg, 1900)^[5]

Potamididae (brackish water snails)

• Cerithidea sp.^[5]

Thiaridae

• Melanoides tuberculata (O. F. Müller, 1774)^[7][4][5]
• Neoradina prasongi^[5]
• Sermyla requetii^[6][5] - synonym: Sermyla tornatela^[4][5]
• Tarebia granifera (Lamarck, 1822)^[6][4][5]
• Thiara scabra^[6][4][5]

Assimineidae

• Assiminea brevicula^[5]
• Assiminea francoisi^[5]
• Assiminea interrupta^[5]
• Assiminea obtusa^[5]
• Assiminea lutea^[5]

Bithyniidae

• Digoniostoma siamensis^[5]
• Parafossarulus striatulus - synonym: Parafossarulus manchouricus^[5]
• Alocinma longicornis^[5]
• Bithynia misella^[5]
• Bithynia fuchsiana^[4][5]
• Bithynia goniomphalos^[5]
• Wattebledia crosseana^[5]

Pomatiopsidae

• Neoprososthenia levayi - synonym: Paraprososthenia levayi^[5]
• Lacunopsis harmandi^[5]
• Tricula ovata^[5]
• Tricula similunaris^[5]
• Vietricula - 11 species^[5]

Stenothyridae

• Stenothyra divalis^[5]
• Stenothyra messageri^[6][4][5]
• Stenothyra monilifera^[5]
• Stenothyra polita^[5]
• Stenothyra cyrtochila^[5]
• Stenothyra ovata^[5]
• Stenothyra conica^[5]
• Stenothyra alba^[5]
• Stenothyra schlickumi Brandt, 1968^[5]

Iravadiidae (marine/brackish)

• Fluvicingula elegantula (A. Adams, 1861)^[5]
• Iravadia cochinchinensis (Bavay & Dautzenberg, 1910)^[5]
• Iravadia ornata Blanford, 1867^[5]
• Iravadia quadrasi (O. Boettger, 1893)^[5]
• Iravadia rohdei (Brandt, 1968)^[5]
• Pseudomerelina mahimensis (Melvill, 1893)^[5]
• Hyala sp.^[5]

Buccinidae

• Clea helena (Meder in Philippi, 1847)^[5]

Lymnaeidae

• Austropeplea viridis (Quoy & Gaimard, 1832)^[8]
• Radix auricularia (Linnaeus, 1758) - synonym: Lymnaea swinhoei^[8]
• Radix rubiginosa (Michelin, 1831)^[8]

Planorbidae

• Polypylis hemisphaerula (Benson, 1842)^[4]
• Gyraulus convexiusculus (Hutton, 1849)^[4]

Land gastropods

Land gastropods in Vietnam include:

Specimens of Cyclophorus phongnhakebangensis (a: holotype, b and c: paratypes; scale bar: 10 mm)

Cyclophoridae

• Chamalycaeus depressus (Bavay et Dautzenberg, 1912)^[9]
• Chamalycaeus fraterculus (Bavay et Dautzenberg, 1900)^[9]
• Chamalycaeus heudei (Bavay et Dautzenberg, 1900)^[9]
• Chamalycaeus paviei (Bavay et Dautzenberg, 1912)^[9]
• Dioryx compactus (Bavay et Dautzenberg, 1900)^[9]
• Dioryx messageri (Bavay et Dautzenberg, 1900)^[9]
• Dioryx vanbuensis (Bavay et Dautzenberg, 1900)^[9]
• Laotia christahemmenae Páll-Gergely, 2014^[9]
• Cyclophorus affinis Theobald, 1858^[10]
• Cyclophorus amoenus (Pfeiffer, 1854)^[10]
• Cyclophorus aquilus (Sowerby, 1843)^[10]
• Cyclophorus aurantiacus pernobilis Gould, 1844^[10]
• Cyclophorus bensoni (Pfeiffer, 1854)^[10]
• Cyclophorus cambodgensis Morlet, 1884^[10][9]
• Cyclophorus cantori (Benson, 1851)^[10]
• Cyclophorus clouthianus Möllendorff, 1882^[10]
• Cyclophorus consociatus Smith, 1893^[10]
• Cyclophorus courbeti Ancey, 1888^[11]
• Cyclophorus courbeti courbeti Ancey, 1888^[10]
• Cyclophorus courbeti leucostoma Dautzenberg et Fischer, 1905^[10]
• Cyclophorus cucphuongensis Oheimb, 2019^[11]
• Cyclophorus cryptomphalus Benson, 1857^[10]
• Cyclophorus dilatatus Heude, 1886^[10]
• Cyclophorus dodrans dodrans Mabille, 1887^[10]
• Cyclophorus dodrans fasciatus Kobelt, 1908^[10]
• Cyclophorus donghoiensis Thach et Huber, 2017^[10]
• Cyclophorus eudeli Smith, 1893^[10]
• Cyclophorus expansus (Pfeiffer, 1853)^[10]
• Cyclophorus fargesianus Heude, 1885^[10]
• Cyclophorus fasciatus Kobelt, 1908^[11]
• Cyclophorus floridus (Pfeiffer, 1854)^[10]
• Cyclophorus fruhstorferi fruhstorferi Möllendorff, 1901^[10]
• Cyclophorus fruhstorferi langsonensis Kobelt, 1908^[10]
• Cyclophorus fulguratus Pfeiffer, 1852^[10]
• Cyclophorus haughtoni Theobald, 1858^[10]
• Cyclophorus herklotsi Martens, 1861^[10]
• Cyclophorus huberi Thach, 2016^[10]
• Cyclophorus ignilabris Möllendorff, 1901^[10]
• Cyclophorus implicatus Bavay et Dautzenberg, 1908^[9]
• Cyclophorus implicatus implicatus Bavay et Dautzenberg, 1908^[10]
• Cyclophorus implicatus kanhoensis Do et Do, 2019^[10]
• Cyclophorus jourdyi Morlet, 1886^[11][10]
• Cyclophorus labiosus (Pfeiffer, 1854)^[10]
• Cyclophorus malayanus (Benson, 1852)^[10]
• Cyclophorus mansuyi Dautzenberg et Fischer, 1908^[10]
• Cyclophorus martensianus Möllendorff, 1874^[10]
• Cyclophorus massiei Morlet, 1891^[10][9]
• Cyclophorus monachus (Morelet, 1866)^[10]
• Cyclophorus muspratti Godwin-Austen et Beddome, 1894^[10]
• Cyclophorus orthostylus Möllendorff, 1898^[10]
• Cyclophorus paracucphuongensis Oheimb, 2019^[11]
• Cyclophorus paviei Morlet, 1884^[10]
• Cyclophorus pfeifferi Reeve, 1861^[10]
• Cyclophorus phongnhakebangensis Oheimb, 2019^[11]
• Cyclophorus polystictus Möllendorff, 1901^[10]
• Cyclophorus pyrostoma Möllendorff, 1882^[10]
• Cyclophorus saturnus Pfeiffer, 1862^[10]
• Cyclophorus siamensis (Sowerby, 1850)^[10]
• Cyclophorus songmaensis Morlet, 1891^[10]
• Cyclophorus speciosus (Philippi, 1847)^[10]
• Cyclophorus subfloridus Ancey, 1888^[11][10]
• Cyclophorus takumisaitoi Hirano, 2019^[11]
• Cyclophorus tamdaoensis Do et Do, 2019^[10]
• Cyclophorus tetrachrous Mabille, 1887^[10]
• Cyclophorus theodori Ancey, 1888^[10]
• Cyclophorus tornatus Morlet, 1892^[10]
• Cyclophorus trouiensis trouiensis Wattebled, 1886^[10]
• Cyclophorus trouiensis bendei Varga, 1972^[10]
• Cyclophorus trouiensis omphalotropis Möllendorff, 1900^[10]
• Cyclophorus unicus Mabille, 1887^[10]
• Cyclophorus volvulus (O.F. Müller, 1774)^[10][9]
• Cyclophorus zebrinus (Benson, 1836)^[10]
• Cyclotus lubricus (Dautzenberg et Fischer, 1908)^[9]
• Japonia fischeri (Morlet, 1886)^[9]
• Japonia scissimargo (Benson, 1856)^[9]
• Platyrhaphe leucacme Möllendorff, 1901^[9]
• Platyraphe sordida (Pfeiffer, 1855)^[9]
• Pterocyclos prestoni Bavay et Dautzenberg, 1909^[9]
• Rhiostoma morleti Dautzenberg et Fischer, 1905^[9]
• Scabrina laciniana (Heude, 1885)^[9][12]
• Scabrina locardi (J. Mabille, 1887)^[12]
• Scabrina tonkiniana (J. Mabille, 1887)^[12]
• Scabrina vanbuensis (Smith, 1896)^[9][12]

Diplommatinidae

• Diplommatina balansai Morlet, 1886^[9]
• Diplommatina clausilioides Bavay et Dautzenberg, 1912^[9]
• Diplommatina demangei Bavay et Dautzenberg, 1912^[9]
• Diplommatina messageri Ancey, 1903^[9]

Pupinidae

• Pupina anceyi Bavay & Dautzenberg, 1899^[13][9]
• Pupina billeti Fischer, 1898^[13]
• Pupina brachysoma Ancey, 1903^[13]
• Pupina dorri Dautzenberg, 1893^[13][9]
• Pupina douvillei Dautzenberg & Fischer, 1905^[13]
• Pupina exclamationis Mabille, 1887^[13][9]
• Pupina flava Möllendorff, 1884^[13]
• Pupina illustris Mabille, 1887^[13]
• Pupina laffonti Ancey, 1899^[13]
• Pupina solidula Möllendorff, 1901^[13]
• Pupina sonlaensis Do, 2017^[13]
• Pupina thaitranbaii Do, 2017^[13]
• Pupina tonkiana Bavay & Dautzenberg, 1899^[13][9]
• Pupina verneaui Dautzenberg & Fischer, 1905^[13][9]
• Pupina vescoi Morelet, 1862^[13]
• Pseudopomatias amoenus Möllendorff, 1885^[9]
• Pseudopomatias maasseni Páll-Gergely et Hunyadi, 2015^[9]
• Pseudopomatias sophiae Páll-Gergely, 2015^[9]
• Pupinella mansuyi (Dautzenberg et Fischer, 1908)^[9]
• Rhaphaulus tonkinensis Páll-Gergely, 2014^[9]
• Streptaulus longituba Páll-Gergely & Gargominy, 2017^[14]

Helicinidae

• Geotrochatella jourdyi Dautzenberg, 1895^[9]

Hydrocenidae

• Georissa decora Möllendorff, 1900^[9]

A shell of Leptacme cuongi

Clausiliidae

• Castanophaedusa fontainei Páll-Gergely & Szekeres, 2017^[15]
• Castanophaedusa huberi (Thach, 2016)^[15]
• Garnieria mouhoti nhuongi Do, 2015^[15]
• Grandinenia ardouiniana (Heude, 1885)^[15]
• Grandinenia tonkinensis (Nordsieck, 2010)^[15]
• Hemiphaedusa babeensis crassitesta Nordsieck, 2011^[16]
• Hemiphaedusa porphyrostoma regina Nordsieck, 2011^[16]
• Hemiphaedusa thatkheana ptychostoma Nordsieck, 2011^[16]
• Hemiphaedusa thatkheana splendida Nordsieck, 2011^[16]
• Leptacme cuongi Maassen & Gittenberger, 2007^[2]
• Leptocochlea sykesi lethrungtongi Grego & Szekeres, 2011^[17]
• Liparophaedusa leptotesta Nordsieck, 2011^[16]
• Megalauchenia soror Nordsieck, 2011^[16]
• Messageriella gargominyi Páll-Gergely & Szekeres, 2017^[15]
• Oospira abstrusa Szekeres, 1970^[15]
• Oospira abstrusa ginkae Grego & Szekeres, 2014^[15]
• Oospira duci Maassen & Gittenberger, 2007^[2]
• Oospira duci khanhi Nordsieck, 2011^[16]
• Oospira haivanensis Chinh & Szekeres, 2019^[18]
• Oospira naggsi Luong & Szekeres, 2014^[15]
• Oospira naggsi naggsi Luong & Szekeres, 2014^[15]
• Oospira naggsi parva Páll-Gergely & Szekeres, 2017^[15]
• Oospira oviformis Nordsieck, 2011^[16]
• Oospira pyknosoma Gittenberger & Vermeulen, 2001^[19]
• Oospira smithi Maassen & Gittenberger, 2007^[2]
• Oospira triptyx Nordsieck, 2011^[16]
• Phaedusa cochinchinensis (Pfeiffer, 1841)^[18]
• Phaedusa lypra pereupleura Nordsieck, 2011^[16]
• Phaedusa micropaviei Nordsieck, 2011^[16]
• Synprosphyma oospiroides Nordsieck, 2011^[16]
• Tropidauchenia bavayi senescens Nordsieck, 2011^[16]
• Tropidauchenia (Euryauchenia) fisheri phasmoides Grego & Szekeres, 2011^[17]
• Tropidauchenia ootanii longicollis Nordsieck, 2011^[16]
• Tropidauchenia palatalis Nordsieck, 2011^[16]

Helicarionidae

• Falsiplecta integripedia Schileyko & Semenyuk, 2018^[20]
• Laocaia attenuata Kuzminykh, 1999^[21]
• Laocaia obesa Kuzminykh, 1999^[21]
• Laocaia simovi Dedov & Schneppat, 2019^[21]

Streptaxidae - 39 species of Streptaxidae are known from Vietnam^[22]

• Discartemon^[22]
• Indoartemon^[22]
• Perrottetia aberrata (Souleyet, 1852)^[22]
• Perrottetia cristatellus (Möllendorff, 1901)^[22]
• Perrottetia daedaleus (Bavay & Dautzenberg, 1908)^[22]
• Perrottetia dugasti (Morlet, 1892)^[22]
• Perrottetia gudei (Fulton, 1915)^[22]
• Perrottetia hongthinhae D.-S. Do, 2017^[22]
• Perrottetia mabillei (Bavay et Dautzenberg, 1903)^[22]
• Perrottetia messageri (Bavay et Dautzenberg, 1908)^[22]
• Perrottetia oppidulum (Bavay et Dautzenberg, 1908)^[22]
• Perrottetia simonianus (Heude, 1890)^[22]
• Stemmatopsis arcuatolabris Do, 2021^[23]
• Stemmatopsis nangphaiensis Do & Do, 2015^[23]
• Stemmatopsis vanhoensis Do & Do, 2015^[23]
• Haploptychius^[22]
• Oophana^[22]
• Elma Adams, 1866^[24][22]
• Huttonella^[22]

Plectopylidae - 16 species are known from Vietnam^[25]

Gudeodiscus messageri

• Gudeodiscus anceyi (Gude, 1901)^[25]
• Gudeodiscus cyrtochilus (Gude, 1909)^[25]
• Gudeodiscus dautzenbergi (Gude, 1901)^[25]
• Gudeodiscus fischeri (Gude, 1901)^[25]
• Gudeodiscus francoisi (Fischer, 1898)^[25]
• Gudeodiscus giardi (Fischer, 1898)^[25]
• Gudeodiscus hemmeni Páll-Gergely & Hunyadi, 2015^[25]
• Gudeodiscus infralevis (Gude, 1908)^[25]
• Gudeodiscus messageri (Gude, 1909)^[25]
• Gudeodiscus phlyarius (Mabille, 1887)^[25]
• Gudeodiscus suprafilaris (Gude, 1908)^[25]
• Gudeodiscus villedaryi (Ancey, 1888)^[25]
• Gudeodiscus emigrans (Möllendorff, 1901)^[25]
• Halongella fruhstorferi (Möllendorff, 1901)^[25]
• Halongella schlumbergeri (Morlet, 1886)^[25]
• Sicradiscus mansuyi (Gude, 1908)^[25]

Philomycidae

• Meghimatium bilineatum (Benson, 1842)^[26]

Ariophantidae

• Hemiplecta jensi Páll-Gergely, 2019^[27]
• Parmarion martensi Simroth, 1894^[26]
• Parmarion pupillaris Humbert, 1864^[26]

Agriolimacidae

• Deroceras laeve (O. F. Müller, 1774)^[28]

Camaenidae

• Amphidromus cambojiensis (Reeve, 1860)^[29]
• Amphidromus ingens Möllendorff, 1900^[29]
• Amphidromus naggsi Thach & Huber, 2014^[29]

Freshwater bivalves

Freshwater bivalves in Vietnam include:

See also

• List of marine molluscs of Vietnam

Lists of molluscs of surrounding countries:

• List of non-marine molluscs of China
• List of non-marine molluscs of Laos
• List of non-marine molluscs of Cambodia

References

This article incorporates CC-BY-3.0 text from the reference^[2]

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27. Barna Páll-Gergely (2019). "A new species of Hemiplecta Albers, 1850 from Vietnam (Gastropoda: Pulmonata: Ariophantidae)". Folia Malacologica 27(2): 85–89. DOI: https://doi.org/10.12657/folmal.027.007
28. Ivailo Kanev Dedov, Ulrich E. Schneppat, Heike Reise, Manh Quang Vu (2020). "First record of an agriolimacid slug in Southeast Asia – Deroceras laeve (O. F. Müller, 1774) (Gastropoda: Pulmonata) recently introduced to the Socialist Republic of Vietnam". Biodiversity Data Journal 8: e59644
29. Thach, Nguyen Ngoc; Huber, Franz (2014). "A new Amphidromus (Gastropoda, Camaenidae) from Vietnam". Basteria. 78 (1–3): 35–37.

Further reading

• Do D. S., Nguyen T. H. T. & Do V. N. (2015). "A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam". Ruthenica 25(4): 117–132. PDF.
• Vermeulen J. J. & Whitten T. (1998). "Land and freshwater molluscs of the karst regions ENE of Haiphong and the Cuc Phuong National Park, northern Vietnam". Report of a survey for the Vietnam Programme of FFI 12 pp. (unpublished report).
• Vermeulen J. J. & Maassen W. J. M. (2003). "The non-marine mollusk fauna of the Pu Luong, Cuc Phuong, Phu Ly, and Ha Long regions in northern Vietnam". Report of a survey for the Vietnam Programme of FFI 35 pp. (unpublished report).