Hapalodectes (literal translation 'soft biter'; from Ancient Greek: ἁπαλόςhapalos ('soft, tender') and δῆκτῆςdêktês ('biter')) is an extinctgenus of otter-like mesonychians from the Late Paleocene to Early Eocene, some 55million years ago. Although the first fossils were found in the Eocene strata of Wyoming, the genus originated in Mongolia, as the oldest species is H. dux, which was found in Late Paleocene strata in the Naran Bulak Formation.[2]
The genus was once suggested to be related to the Archaeoceti, such as Pakicetus, due to numerous similarities between the skull and tooth anatomies of the two genera. Now, however, Hapalodectes and other mesonychians are thought to be related to basalartiodactyls, while the Archaeoceti are now determined to be descended from more derived artiodactyls, like Indohyus, which are related to hippopotamuses and anthracotheres.[citation needed]
H. dux possessed distinct conules on its upper molars.[3]