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Desmodium
	Desmodium heterocarpon
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Rosids
Order:	Fabales
Family:	Fabaceae
Subfamily:	Faboideae
Tribe:	Desmodieae
Subtribe:	Desmodiinae
Genus:	Desmodium; Desv. (1813), nom. cons.
Species
	Many, see text
Synonyms
	Cyclomorium Walp. (1843); Edusaron Medik. (1787), nom. superfl.; Meibomia Heist. ex Fabr. (1759), nom. rej.; Nephromeria (Benth.) Schindl. (1924); Nicolsonia DC. (1825); Nissoloides M.E.Jones (1933); Sagotia Duchass. & Walp. (1851); Tropitoma Raf. (1837);

Desmodium is a genus of plants in the legume family Fabaceae, sometimes called tick-trefoil, tick clover, hitch hikers or beggar lice.^[2] There are dozens of species and the delimitation of the genus has shifted much over time. Species are distributed widely – from Quebec to northern Argentina in the Americas, across northern and southern tropical Africa, in the southern Arabian Peninsula, in Myanmar and Thailand, New Guinea, and northern and eastern Australia.^[1]

Description

These are mostly inconspicuous plants; few have bright or large flowers. Though some can become sizeable plants, most are herbs or small shrubs. Their fruit are loments, meaning each seed is dispersed individually enclosed in its segment. This makes them tenacious plants and some species are considered weeds in places.

Uses

Several Desmodium species release organic compounds, aerially and into the soil, which make them useful for agriculture: Allelopathic compounds are used there via push-pull technology. For this Desmodium heterocarpon, Desmodium intortum, and Desmodium uncinatum are inter-cropped in maize and sorghum fields to suppress witchweeds, including Asiatic witchweed (Striga asiatica) and purple witchweed (S. hermonthica) and to repel Chilo partellus, a stem-boring grass moth.^[3] Insects (including pests) are likewise repelled by high amounts of antixenotic allomones produced by Desmodium.

A blue dye is obtained from Desmodium incanum.

Tick-trefoils in agriculture can also be used as living mulch and as green manure, as they improve soil fertility via nitrogen fixation.

Most also make good fodder for animals including bobwhite, turkey, grouse, deer, cattle and goats.^[3]^[4]^[5]

Wild-living, non-farmed deer appear to rely on Desmodium species in certain areas, particularly during the more stressful summer months.

The caterpillars of the lesser grass blue (Zizina otis) and the two-barred flasher (Astraptes fulgerator) feed on tick-trefoils.

Alkaloids

Some Desmodium species have formerly been known to contain high amounts of tryptamine alkaloids, but many of the tryptamine-containing species have since been transferred to other genera.^[3]

Taxonomy and systematics

The taxonomy and systematics of the many dozens of Desmodium species are confusing and unresolved. Related genera such as Codariocalyx, Hylodesmum, Lespedeza, Ohwia, and Phyllodium were and sometimes still are included in Desmodium.^[6]

Taxonomic authorities commonly disagree about the naming and placement of species. For example, Desmodium spirale as described by August Grisebach might refer to a distinct species, but its validity is doubtful. The "Desmodium spirale" of other authorities may refer to D. neomexicanum, D. ospriostreblum, or D. procumbens. Similarly, the plant originally described as D. podocarpum by A. P. de Candolle is Hylodesmum podocarpum today, but "Desmodium podocarpum" might also refer to D. hookerianum or Hylodesmum laxum, depending on the taxonomic authority.^[6]

Selected species

Species include:^[1]^[6]^[7]

Desmodium acanthocladum F.Muell.
Desmodium canadense (L.) DC. – showy tick-trefoil, Canadian tick-trefoil
Desmodium canescens (L.) DC. – hoary tick-trefoil
Desmodium ciliare (Muhl.) DC. – hairy small-leaved tick-trefoil
Desmodium cuspidatum (Muhl.) Loudon – toothed tick-trefoil, large-bracted tick-trefoil
Desmodium fernaldii B.G.Schub.
Desmodium glabellum (Michx.) DC.
Desmodium heterocarpon (Michx.) DC. – Asian tick-trefoil
Desmodium × humifusum (Muhl. ex Bigelow) Beck
Desmodium illinoense A.Gray – Illinois tick-trefoil
Desmodium incanum (Sw.) DC. – creeping beggarweed, Spanish tick-trefoil, Kaimi clover
Desmodium intortum Greenleaf desmodium, kuru vine, beggarlice, tick clovers
Desmodium lineatum (Michx.) DC. – linear-leaved tick-trefoil
Desmodium marilandicum (L.) DC. – smooth small-leaved tick-trefoil
Desmodium molliculum (Kunth) DC.
Desmodium ospriostreblum Chiov.
Desmodium paniculatum (L.) DC. – panicled tick-trefoil
Desmodium perplexum B.G.Schub. – perplexed tick-trefoil
Desmodium rotundifolium DC. – round-leaved tick-trefoil, dollar leaf
Desmodium sessilifolium (Torr. ex M.A.Curtis) Torr. & A.Gray
Desmodium tortuosum (Sw.) DC.
Desmodium triflorum (L.) DC.
Desmodium tweedyi Britton – Tweedy's tick-trefoil
Desmodium uncinatum (Jacq.) DC.– silver-leaved tick-trefoil, silverleaf
Desmodium varians (Labill.) G.Don

Formerly placed here

Codariocalyx motorius – telegraph plant (as D. gyrans, D. motorium, D. roylei)
Dendrolobium triangulare (as D. Desmodium umbellatum Moritz.)
Dendrolobium umbellatum (as D. umbellatum (L.) Benth. )
Hylodesmum laxum (as D. laxum DC.)
- Hylodesmum laxum ssp. laxum (as D. austro-japonense, D. bambusetorum, D. gardneri auct. non Benth., D. laxiflorum sensu Miq., D. laxum var. kiusianum, D. laxum ssp. laxum, D. podocarpum auct. non DC. non Hook. & Arn., D. podocarpum DC. var. gardneri sensu Bedd., D. podocarpum DC. var. laxum)
Hylodesmum leptopus (as D. gardneri Benth., D. laxum auct. non DC., D. laxum ssp. leptopus, D. leptopus, D. tashiroi)
Hylodesmum podocarpum (as D. podocarpum DC., D. podocarpum DC. var. indicum, D. podocarpum DC. var. japonicum)
- Hylodesmum podocarpum ssp. oxyphyllum (as D. fallax var. mandshuricum, D. japonicum, D. mandshuricum, D. oxyphyllum DC., D. podocarpum DC. var. mandshuricum, D. podocarpum DC. ssp./var. oxyphyllum, D. podocarpum DC. var. polyphyllum, D. podocarpum DC. var. typicum, D. racemosum)
Lespedeza thunbergii (as D. formosum, D. thunbergii)