He was born in Zhenhai, Zhejiang province, on October 10, 1903.He found the department of biology, Zhejiang University in 1929, and work there for 20 years. He was the oldest member of both the Academia Sinica and the Chinese Academy of Sciences at the time of his death. He was the founder, the first chief director and honorary director of the Institute of Biophysics, Chinese Academy of Sciences.