During the long vacation of 1889 and again in the following year he proceeded to Messina at the instance of Lankester and there carried out the investigations of Amphioxus which formed the subject matter of his first published papers. In 1891 he spent some months at Freiburg under Weismann and later on spent a year at the Marine Stations of Plymouth and Naples, working chiefly on the development of the Ascidians.[2]
After returning from the Marine Station (Stazione Zoologica), Naples, he became a fellow at University College London. From 1892 to 1894 he was a biological tutor at Columbia University. From 1894 to 1899 he held the Balfour Studentship of Cambridge, during which he went to the East Indies to investigate the embryology of the pearly nautilus. From 1899 to 1901 he was a lecturer in biology at Guy's Hospital. In 1902 he was elected F.R.S. From 1902 to 1909 he was the director of the Colombo Museum and the editor of Spolia Zeylanica.[3] In 1907 he was a marine biologist for the Ceylon Government. In 1910 he became the Strathcona Professor at McGill University in Montreal[4] and retired there as professor emeritus in 1932. In 1902 in Hendon, Middlesex, he married Emily Constance Bowd, and after his retirement in 1932 they lived in a cottage about an hour's drive from Montreal.
^Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). "Willey". The Eponym Dictionary of Reptiles. Baltimore: Johns Hopkins University Press. p. 286. ISBN9781421401355.